Nancy H. Ruddle, Ph.D.

Nancy H. Ruddle, Ph.D. ’68, professor emerita and senior research associate in the Division of Epidemiology of Microbial Diseases in the School of Public Health, received the 2009 Lifetime Achievement Award from the International Cytokine Society at its annual meeting in Lisbon, Portugal, in October. The award is given annually to a scientist who has contributed to research on cytokines, proteins released by cells within the immune system that are critical to regulating the body’s immune response. Ruddle’s research, which began during her years as a graduate student in microbiology at Yale, focuses on lymphotoxin and other cytokines involved in autoimmune diseases and lymphoid organ development.