Faculty

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Jeffrey Bender, MD

Robert I. Levy Professor of Medicine (Cardiology) and Professor of Immunobiology

Associate Chief, Cardiovascular Medicine, Internal Medicine

Director, Yale Cardiovascular Research Center, Internal Medicine

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Marcus Bosenberg, MD, PhD

Professor of Dermatology, Pathology, and Immunobiology

Marcus Bosenberg M.D., Ph.D., is a physician scientist who directs a leading melanoma research laboratory, is Co-Leader of the Genomics, Genetics and Epigenetics Program of the Yale Cancer Center, Co-Director of the Yale SPORE in Skin Cancer, and is a practicing dermatopathologist at Yale Dermatopathology through Yale Medicine.

In his research, Dr. Bosenberg studies the genetics and cellular changes that result in melanoma, the leading cause of skin cancer deaths. His laboratory has developed...

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Alfred Bothwell, PhD

Professor of Immunobiology

Member of HTI and VBT

Al Bothwell graduated with an A.B. from Washington University in 1971, got a PhD from Yale in Sidney Altman’s lab in 1975 and then did a postdoc with David Baltimore at MIT where he established the genetic basis of the anti-NP idiotypic antibody response. He has been on the Immunobiology faculty at the Yale Medical School since 1982. He continued studies of B cell antibody diversity and memory and then worked on T cell receptor structure/function and signaling. He also developed the molecular...

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Grace Chen, PhD

Assistant Professor

Grace Chen received her undergraduate training in the College of Chemistry at UC Berkeley. She attended Harvard University for her PhD where she worked in David Liu's laboratory to discover and characterize novel RNA modifications. Her postdoctoral research was at Stanford University in Howard Chang's group, where she investigated circular RNA immunity. Grace Chen joined Yale University as a faculty in the Department of Immunobiology in 2019. Her research focuses on the functions and...

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Lieping Chen, MD, PhD

United Technologies Corporation Professor in Cancer Research and Professor of Immunobiology, of Dermatology and of Medicine (Medical Oncology)

Co-Director, Cancer Immunology Program at Yale Cancer Center

Lieping Chen studies immune cell communications via cell surface protein-protein interactions. He is also interested in translating laboratory findings to treat human diseases including cancer, autoimmune diseases and infection. 

In 1992, Dr. Chen showed the first proof-of-concept study that the B7-CD28 family molecules could be the targets for cancer immunotherapy. This study inspires subsequent studies targeting the B7-CD28 family molecules for the treatment of human cancer.

In 1999, Dr. Chen,...

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Joseph Craft, MD

Paul B. Beeson Professor of Medicine (Rheumatology) and Professor of Immunobiology

Paul B. Beeson Professor of Medicine

Program Director, Investigative Medicine

Dr. Joseph Craft is Paul B. Beeson Professor of Medicine and Professor of Immunobiology at the Yale University School of Medicine. He received his degree in chemistry as a Phi Beta Kappa graduate of University of North Carolina at Chapel Hill, and is an Alpha Omega Alpha graduate of the University of North Carolina School of Medicine. Dr. Craft did postgraduate training in medicine and in rheumatology and immunology at Yale, and has been on the faculty at that institution since 1985. At Yale,...

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Peter Cresswell, PhD, FRS

Eugene Higgins Professor of Immunobiology and Professor of Cell Biology

Dr. Cresswell is the Eugene Higgins Professor of Immunobiology and Professor of Cell Biology and Dermatology at Yale University School of Medicine.



He received his B.S. degree in chemistry, his M.S. degree in microbiology from the University of Newcastle Upon Tyne, U.K., and his Ph.D. degree in biochemistry and immunology from London University. His postdoctoral training was completed at Harvard University with Jack Strominger.



Before assuming his position at Yale, Dr. Cresswell was Chief of the...

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Vishwa Deep Dixit, DVM, PhD

Waldemar Von Zedtwitz Professor of Comparative Medicine and of Immunobiology

Vishwa Deep Dixit completed Bachelor and Master of Veterinary Sciences in HAU, Hisar India. He received German Academic Exchange Service fellowship to conduct PhD research in Germany. He completed PhD coursework in HAU and Research Work in University of Hannover in Year 2000. He conducted postdoctoral research training in NIH. He joined Pennington Biomedical Research Center as an Assistant Professor in 2006 and moved to Yale as Professor of Comparative Medicine and Immunobiology in 2013. ...

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Stephanie Eisenbarth, MD, PhD

Associate Professor of Laboratory Medicine, of Immunobiology and of Medicine (Immunology)

Associate Chair of Research, Laboratory Medicine

Assistant Director of Clinical Pathology Residency Program

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Richard Flavell, PhD, FRS

Sterling Professor of Immunobiology

Investigator, Howard Hughes Medical Institute

Dr. Flavell is Sterling Professor of Immunobiology at Yale University School of Medicine, and an Investigator of the Howard Hughes Medical Institute. He received his B.Sc. (Honors) in 1967 and Ph.D. in 1970 in biochemistry from the University of Hull, England, and performed postdoctoral work in Amsterdam (1970-72) with Piet Borst and in Zurich (1972-73) with Charles Weissmann. Before accepting his current position in 1988, Dr. Flavell was first Assistant Professor (equivalent) at the University...

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Ellen F Foxman, MD, PhD

Assistant Professor of Laboratory Medicine and Immunobiology

Dr. Foxman's research interest is understanding the natural mechanisms that protect the airway from respiratory viruses. Recent evidence shows that respiratory viruses enter the airway much more frequently than they cause illness. Current projects focus on (1) identifying the antiviral defense mechanisms of the airway epithelium, (2) understanding how the environment influences antiviral defense and tips the balance between health and disease upon exposure to a virus, and (3) using biomarkers...

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Ann Haberman, PhD

Associate Professor of Immunobiology

Director, In Vivo Imaging Facility

Director, Flow Cytometry Facility

Ph.D., University of Pennsylvania (1992)

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David A. Hafler, MD, FANA

William S. and Lois Stiles Edgerly Professor of Neurology and Professor of Immunobiology

Chair, Department of Neurology

Neurologist-in-Chief, Yale New Haven Hospital

Dr. Hafler is the William S. and Lois Stiles Edgerly Professor and Chairman Department of Neurology, Yale School of Medicine and is the Neurologist-in-Chief of the Yale-New Haven Hospital. He graduated magna cum laude in 1974 from Emory University with combined B.S. and M.Sc. degrees in biochemistry, and the University of Miami School of Medicine in 1978. He then completed his internship in internal medicine at Johns Hopkins followed by a neurology residency at Cornell Medical Center-New York...

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Kevan Herold, MD

Professor of Immunobiology and of Medicine (Endocrinology)

My background and research are in translational immunology. I am interested in understanding the basis for autoimmune diseases and developing new therapies based on our understanding of disease mechanisms. My focus has largely been in the field of autoimmune Type 1 diabetes. The work encompasses basic laboratory work understanding the regulation of autoreactive T cells to clinical trials that involve novel therapeutics. As part of these studies I have also been very interested in analysis of...

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Akiko Iwasaki, PhD

Waldemar Von Zedtwitz Professor of Immunobiology and Molecular, Cellular and Developmental Biology

Professor of Molecular Cellular and Developmental Biology

Investigator, Howard Hughes Medical Institute

Akiko Iwasaki received her Ph.D. from the University of Toronto (Canada) in 1998, and her postdoctoral training from the National Institutes of Health (USA) (1998-2000). She joined Yale University (USA) as a faculty in 2000, and currently is an Investigator of the HHMI and Waldemar Von Zedtwitz Professor of Department of Immunobiology, and of Department of Molecular Cellular and Developmental Biology. Akiko Iwasaki’s research focuses on the mechanisms of immune defense against viruses at the...

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Nikhil Joshi, PhD

Assistant Professor

My laboratory uses intricate tumor models and advanced approaches to investigate immune cell interactions with developing tumors. My goal is to determine mechanistically why these interactions do not lead to more potent anti-tumor responses and to identify entry points for modulating these interactions through genetic manipulation and therapeutic intervention. My previous studies have focused on using established complex mouse models to investigate how subtypes of T cells in the tumor...

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Paula Kavathas, PhD

Professor of Laboratory Medicine, of Immunobiology and of Molecular, Cellular, and Developmental Biology

Associate Chair for Academic Affairs, Laboratory Medicine - Education

Institutional Leader CIRTL Network

Chair, Women's Faculty Forum (2013-2017)

Dr. Kavathas graduated with a B.A. in American Institutions, from the University of Wisconsin, writing her thesis on the role of Science in America in the 1960s. She obtained her Ph.D. in Genetics from the Department of Genetics, founded in 1921 as the first Genetics Department in the country. At Wisconsin with Dr. Robert DeMars on genetic analysis of the MHC region. As a postdoctoral fellowship at Stanford University with Dr. Leonard Herzenberg she developed a novel approach for cloning genes...

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Steven Kleinstein, PhD

Professor

Dr. Steven Kleinstein is a computational immunologist with a combination of "big data" analysis and immunology domain expertise. His research interests include both developing new computational methods and applying these methods to study human immune responses. Dr. Kleinstein received a B.A.S. in Computer Science from the University of Pennsylvania and a Ph.D. in Computer Science from Princeton University. He is currently an Associate Professor of Pathology (with a secondary appointment in...

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Martin Kriegel, MD, PhD

Associate Professor Adjunct

Dr. Kriegel is an Assistant Professor of Immunobiology and of Medicine (Rheumatology) at Yale School of Medicine. In 2001, he received his MD/PhD equivalent at the Friedrich-Alexander University of Erlangen, Germany, followed by the German Medical Licensure in 2002. From 2003 to 2006, he performed postdoctoral training in immunology at Yale with Dr. Richard Flavell before completing a medicine residency and rheumatology fellowship at Harvard (Beth Israel Deaconess Medical Center and Brigham &...

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Carrie L Lucas, PhD

Assistant Professor of Immunobiology

Dr. Carrie L. Lucas received her PhD from Harvard Medical School and her postdoctoral training from the National Institutes of Health, NIAID. Her laboratory investigates signaling in T cells from healthy people and patients with inherited immune disorders to dissect pathways critical for adaptive immunity. A major focus of her work has been on phosphoinositide 3-kinase (PI3K) signaling and mechanisms of disease in immunodeficient patients with activating mutations in PI3K subunits.

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John D MacMicking, PhD

Associate Professor of Microbial Pathogenesis and of Immunobiology

Member, Yale Systems Biology Institute

Investigator, Howard Hughes Medical Institute

Dr. MacMicking trained in synthetic organic chemistry and biochemistry at the Australian National University (B.Sc, 1st Class Honours) where he conducted thesis work within the Department of Immunology & Cell Biology formerly headed by 1996 Nobel Laureate, Peter Doherty, at the the John Curtin School of Medical Research.

He then pursued Ph.D studies within the Immunology program at Sloan-Kettering Institute-Cornell University in New York City (with Carl Nathan) before post-doctoral work as an...

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Ruslan Medzhitov, PhD

Sterling Professor of Immunobiology

Investigator, Howard Hughes Medical Institute

Medzhitov was born in Tashkent, Uzbekistan, and earned a B.S. at Tashkent State University before going on to pursue a Ph.D. in biochemistry at Moscow University in 1990. He performed his postdoctoral studies with the late Charles A. Janeway Jr. at Yale University Medical School.

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Eric Meffre, PhD

Associate Professor of Immunobiology and of Medicine (Immunology)

My work focuses on the etiology of autoimmune diseases affecting millions of individuals in the world by identifying molecules and pathways involved in the establishment of B-cell tolerance through the investigation of rare patients with primary immunodeficiency (PID), enrolled at Yale and through an international network.

Patients with PID provide opportunities to study the impact of specific gene defects on the regulation of B-cell tolerance and the removal of developing autoreactive B cells...

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Kevin O'Connor, PhD

Associate Professor of Neurology

Dr. Kevin C. O’Connor is an Associate Professor of Neurology and Immunobiology at Yale School of Medicine. He earned a Bachelor of Science degree in Chemistry from the University of Massachusetts at Amherst and his Ph.D. in Biochemistry at Tufts Medical School. He took his post-doctoral training in Immunology at Harvard Medical School where he also spent several years on the faculty as an Assistant Professor. His investigative interests are in human translational immunology and neurology. He...

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Noah Palm, PhD

Assistant Professor of Immunobiology

Noah W. Palm is an Assistant Professor of Immunobiology and a member of the Human and Translational Immunology Program at the Yale University School of Medicine. His laboratory focuses on illuminating the myriad interactions between the immune system and the gut microbiota in health and disease. Dr. Palm performed his doctoral work with Ruslan Medzhitov and his postdoctoral work with Richard Flavell, both at Yale University.

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Joao Pereira, PhD

Associate Professor of Immunobiology

Dr. João P. Pereira is an associate professor of Immunobiology, member of the Yale Cancer Center and of the Yale Stem Cell Center.  He studied Microbiology at Universidade Católica Portuguesa (BSc, 1996), Biotechnology at DeMontfort University (MSc, 1997), and did PhD studies (Universidade do Porto, 2004) in Immunology at the Gulbenkian Institute (Oeiras) and the Pasteur Institute (Paris) mentored by Paulo Vieira. In 2005 Dr. Pereira joined the laboratory of Dr. Jason Cyster (UCSF) as a Howard...

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Jordan Pober, MD, PhD

Bayer Professor of Translational Medicine and Professor of Immunobiology, Pathology and Dermatology

Director, Human and Translational Immunology Program

Vice-Chair, Dept. of Immunobiology for the Section of Human and Translational Immunology

Dr. Pober was born in Brooklyn, New York in 1949 and grew up in the New York City metropolitan area. He attended Haverford College, graduating summa cum laude in 1971 with high honors in Biology, Chemistry and History. He was admitted to Yale’s Medical Scientist Training Program, receiving his MD and his PhD in Molecular Biophysics and Biochemistry with Prof. Lubert Stryer in 1977. He completed his first year of pathology residency at Yale-New Haven Hospital in 1978, was a post-doctoral fellow...

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Aaron Ring, MD/PhD

Assistant Professor of Immunobiology

Aaron Ring received his undergraduate training at Yale University and entered the Stanford Medical Scientist Training Program for his MD and PhD degrees. At Stanford, he worked in the laboratories of K. Christopher Garcia and Irving Weissman to use structure-based protein engineering to develop new cytokine and immune checkpoint therapies for cancer. He additionally developed novel methodologies in protein engineering to create biologic agents against challenging targets such as G protein...

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Carla Rothlin, PhD

Professor

Associate Professor of Pharmacology

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Craig Roy, PhD

Waldemar Von Zedtwitz Professor of Microbial Pathogenesis and of Immunobiology

Vice-Chair, Department of Microbial Pathogenesis

Craig Roy received his B.S. from Michigan State University in 1985 and earned his Ph.D. in Microbiology and Immunology at Stanford University in 1991 in the laboratory of Dr. Stanley Falkow. After completing a postdoctoral fellowship with Dr. Ralph Isberg in the Department of Molecular Microbiology at Tufts University School of Medicine in 1996, he was appointed as an Assistant Professor in the Department of Molecular Genetics and Microbiology at Stony Brook University. Dr. Roy became a...

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David G. Schatz, PhD

Waldemar Von Zedtwitz Professor of Immunobiology and Professor of Molecular Biophysics and Biochemistry

Chair of Immunobiology

Dr. Schatz has made fundamental contributions to our understanding of the mechanisms that assemble and diversify antigen receptor genes that encode antibodies and T cell receptors.  He is best known for the discovery of RAG1 and RAG2, subsequent biochemical insights into RAG function and evolutionary origins, and the discovery of two distinct levels of regulation of somatic hypermutation.

Schatz has co-authored over 160 articles, many in prestigious journals, and has been the recipient of...

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Craig B. Wilen, MD, PhD

Assistant Professor of Laboratory Medicine and of Immunobiology

Dr. Wilen received his A.B in Biology and Economics at Washington University in St. Louis, his MD and PhD from the University of Pennsylvania. His residency training was in clinical pathology at Barnes-Jewish Hospital in St. Louis, MO. His postdoctoral studies were conducted in the laboratory of Herbert "Skip" Virgin in the Department of Pathology & Immunology at Washington University School of Medicine where he studied the pathogenesis of norovirus, the leading cause of acute gastroenteritis...

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Research Scientists

Paul Clark, PhD

Associate Research Scienist

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Elizabeth Eynon, PhD

Research Scientist in Immunobiology

Chair, IACUC

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Nancy Kirkiles-Smith, PhD

Associate Research Scientist in Immunobiology

Laboratory Manager

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Elizabeth Kopp, PhD, BA

Associate Research Scientist in Immunobiology

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Manolis Roulis, PhD

Associate Research Scientist in Immunobiology

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Rashu Seth, PhD

Associate Research Scientist in Immunobiology

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Zuzana Tobiasova

Associate Research Scientist

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Linghua Zheng, PhD

Associate Research Scientist in Immunobiology

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Gloria Zhu, PhD

Associate Research Scientist in Immunobiology

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