Yale Combined Program in the Biological and Biomedical Sciences (BBS)

The Yale Combined Program in the Biological and Biomedical Sciences (BBS) is a doctoral program that enables you to take advantage of all of the resources found at a modern research university. Everything Yale has to offer – faculty, facilities, and campuses - is here in one comprehensive, interdisciplinary graduate program. Explore the 8 Tracks below and the rest of our site to learn more about Yale’s research and training opportunities.

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Biochemistry, Quantitative Biology, Biophysics and Structural Biology

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Computational Biology and Bioinformatics

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Immunology

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Microbiology

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Molecular Cell Biology, Genetics, and Development

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Molecular Medicine, Pharmacology, and Physiology

Neuroscience

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Plant Molecular Biology

BBS has no boundaries, either departmental or geographical. Upon admission to Yale you will have access to courses, seminars, and faculty labs in every department. Because Yale is unusual among major universities in having its main campus and medical school within walking distance of one another, you can easily participate in research activities and courses on the main University campus as well as at the world-renowned Yale School of Medicine. Moreover, Yale’s brand new West Campus is connected to the rest of the university by non-stop shuttle service. Yale in its entirety will be yours to explore.

When you complete your Ph.D. at Yale, you will have received a thorough, interdisciplinary, cutting edge education, and developed key career skills that will leave you extremely well prepared for careers in academia, industry, business, and the many other exciting paths our graduates pursue.

Apply to the BBS Program

Special Initiatives

Chinese Scholarship Council-Yale World Scholars Program

Medical Research Scholars Program (MRSP)

Integrated Graduate Program in Physics, Engineering, and Biology (IGPPEB)

Cooperative MD/PhD Program between UPR and Yale

Yale Cancer Biology Training Program

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