Skip to Main Content

Ryan MacLeod, MD, PhD

Clinical Fellow

Contact Information

Ryan MacLeod, MD, PhD


I completed my BS in Biological Sciences at the University of Arkansas BS followed by MD and PhD in Interdisciplinary Biological Sciences at the University of Arkansas for Medical Sciences. My specialty interest is in endocrinology with a research interest in Bone and Skeletal biology. My hobbies include cooking, soccer, playing with our 3 dogs.

Yale is such an amazing place to work. From your co-residents to fellows to attendings and leadership, everyone has been so genuinely nice and helpful. You never feel alone, and everyone wants you to succeed. When you put that kind of environment together with the great minds and resources that Yale offers, there is so much opportunity for you to become a successful Physician Scientist.

Selected articles:

MacLeod RS, Cawley KM, Gubrij I, Nookaew I, Onal M, O’Brien CA. Effective CRISPR interference of an endogenous gene via a single transgene in mice. Scientific Reports. 9, 17312 (2019).

MacLeod RS, Meyer MB, Xiong J, Cawley KM, Liu Y, Onal M, Benkusky NA, Thostenson JD, Pike JW, O’Brien CA. Deletion of a putative promoter-proximal Tnfsf11 regulatory region in mice does not alter bone mass or Tnfsf11 expression in vivo. PLOS ONE. 16(5): e0250974 (2021).

Education & Training

  • Resident Physician
    Yale New Haven Hospital (2023)
  • MD
    University of Arkansas for Medical Sciences (2021)
  • PhD
    University of Arkansas for Medical Sciences, Interdisciplinary Biomedical Sciences (2021)
  • BS
    University of Arkansas, Biological Sciences (2014)

Departments & Organizations