Entering Classes Information

Our students come from a variety of backgrounds and interests as this picture of our most recent MD-PhD class shows.

Entering Class - 2017

Fang, Calvin

Calvin Fang

  • Year:  1
  • Mentor Information :  Barbara Kazmierczak 
  • Research interest: Generally, neuroscience from genes to cognition. Specifically, the development and function of the many various cell types in circuits underlying perception and behavior
  • Education:  Cornell University, B.S., 2015 (Biology)
  • A little about me : Born in NYC, I have lived up and down the East Coast (including in nearby Branford during elementary school.) Since graduating, I have been spending time in lab at the NIH, paddling with DC Dragon Boat Club, playing table top and board games, and reading as many books as I can before school starts again.
  • Publications :
Feizi, Alborz

Alborz Feizi

  • Year:  1
  • Mentor Information :  Barbara Kazmierczak 
  • Research interest:
  • Education : University of California-Los Angeles, B.S., 2016 (Bioengineering) 
  • A little about me :
  • Publications :

Elsie Gonzalez-Hurtado

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Cellular & Organismal Metabolism, Immunology, Immunometabolism.
  • Education:  University of California-Riverside, B.S., 2015 (Biochemistry) 
  • A little about me:  I was born and raised in the sunny city of Long Beach, CA. Prior to joining the MD/PhD program at Yale, I spent some time living in Baltimore, MD, where I worked as a post-baccalaureate research fellowat the Johns Hopkins School of Medicine. In my spare time I enjoy drawing and painting, yoga, photography, podcasts, cheering on the Golden State Warriors, and drinking coffee while reading a good book.
  • Publications:

Justin Goodwin

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Cancer Metabolism, Infectious Disease Epidemiology
  • Education:  University of Texas at Dallas, B.S., 2015 (Biology), M.S. 2016 (Biotechnology)
  • A little about me:  I like to listen to audiobooks in the lab or read outside of lab. I also enjoy PC gaming, tabletop RPGs, olympic weightlifting, and powerlifting.
  • PublicationsGoodwin J., Neugent M., Lee S., Choe J., Choi H., Jenkins D., Ruthenborg R., Robinson M., Jeong J., Wake M., Abe H., Takeda N., Endo H., Inoue M., Xuan Z., Yoo H., Chen M., Ahn J., Minna J., Helke K., Singh P., Shackelford D. & Kim J. The distinct metabolic phenotype of lung squamous cell carcinoma defines selective vulnerability to glycolytic inhibition. Nature Communications 8, 15503 (2017). DOI: 10.1038/ncomms15503 
  • Oh E., Kim J., Kim J., Kim S., Lee J., Hong, S., Goodwin J., Ruthenborg R., Jung M., Lee H., Lee C., Park E., Kim C. & Park H. NQO1 inhibits proteasome-mediated degradation of HIF-1α. Nature Communications 7, 13593 (2016). DOI: 10.1038/ncomms13593
Hubbard - NEW

Brandon Hubbard

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research:  Understanding the role of metabolism in physiology and disease
  • Education:  Oklahoma State University, B.S., 2015 (Physiology, Psychology)
  • A little about me: I grew up in rural Oklahoma, where my family has lived and farmed since the Land Run of 1889. After attending college at Oklahoma State, I moved to Washington, DC, and worked as a post-baccalaureate research fellow at the NIH for two years. I enjoy tennis, baseball, disc golf, hiking, and mountain biking. I'm also a percussionist and avid music lover.
  • Publications:

Jillian Jaycox

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Immunology
  • Education:  Carnegie Mellon University, B.S., 2016 (biological Sciences); University of Cambridge, M. Phil Medical Science 
  • A little about me:  A little about me: I grew up in southeastern Pennsylvania and then headed to the west side of the state for my undergraduate at Carnegie Mellon. While completing my degree, I ran on the CMU cross country and track teams and developed an interest in autoimmunity through my research at University of Pittsburgh. My scientific interests led me to the University of Cambridge, where I completed a research masters degree focused on B cell biology, while also spending plenty of time rowing, playing croquet, and drinking tea. Outside of science, I love to run, go hiking and backpacking, play guitar, watch documentaries, read nonfiction science and history books, and work with animals (especially dogs and horses). 
  • Publications:

Jonathan Klein

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Infectious Disease, Immunology, Host-Pathogen Interactions, Oncolytic Viruses
  • Education:  University of California-Davis, B.S., 2012 (Biochemistry; Minor: Neuroscience) 
  • A little about me:  I was born and raised in Wine Country in Northern California. I graduated from the University of California, Davis in 2012 and afterwards spent time in the laboratory of Leor Weinberger at UCSF and the Gladstone Institutes studying viral gene circuitry and the mechanisms driving pathogen fate-decisions. I joined the Yale School of Medicine in 2017 as an MD/PhD student with interests in both immunobiology and infectious disease. When I’m not studying or in the laboratory, I love going hiking, camping, and watching terrible 80’s action movies.
  • Publications:

Valentyna Kostiuk

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Cancer biology: gene expression regulation, epigenetics and immunology.
  • Education:  CUNY Hunter College, B.A., 2016 (Biological Sciences) 
  • A little about me:  I was born and raised in Ukraine but attended college in NYC. In my free time I enjoy cooking, reading, coffee, listening to music and long walks.
  • Publications:
Li, Dan

Dan Li

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research Interest: Developmental Biology
  • Education:  Harvard University, B.A., 2015 (Chemical and Physical Biology) 
  • A little about me: I was born and raised in a beachtown called Dalian, China. I moved to the States when I was fifteen years old. I have been moving all over the country ever since -- four years in Portland, four years in Boston, and two years in Dallas. During college, I worked on a project dissecting the  hemoglobin-switch with the hope to find a treatment for the sickle cell disease. In my spare time, I enjoy meditating, reading, jogging along the beach, and working on a nonprofit called Education without Barriers.
  • Publications:
Lin, Kingson

Kingson Lin

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Drug design and synthesis, High-throughput screening, Chemical biology, Mechanisms of drug action.
  • Education:  University of Pennsylvania, B.A., 2017 (Biochemistry); M.S., Chemistry
  • A little about me:  I was born in New York, grew up in China, and came back to the U.S. when I was 5 to start school. I'm a huge chemistry buff and my hobbies include Marvel movies, cooking, tennis, and traveling. I hope to one day become a practicing physician-scientist and corgi owner.
  • Publications:
Lynn, Anna

Anna Lynn

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Targeted nanoparticle drug delivery to the brain
  • Education:  University of Kansas, B.S., 2017 (Chemical Engineering) 
  • A little about me: I was born in China and raised in Kansas, where I went to college and became a devoted Jayhawk basketball fan! In my free time, I enjoy ballet, taekwondo, reading, playing guitar, and spending time with family and friends.
  • Publications:

Neil Savalia

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research:  Brain imaging and computational neuroscience
  • Education:  Washington University, B.A., 2013 (Biology, MB&B, Philosophy-Neuroscience-Psychology) 
  • A little about me:  I grew up in West Orange, NJ, and spent several years in St. Louis, MO for college and in Dallas, TX as a research manager (2013-2017). I joined the MD/PhD Program at the Yale School of Medicine in 2017 with an interest in using novel brain imaging techniques to better understand neuropsychiatric disorders. In my spare time I can be found hanging out in the mountains, experimenting on guitar, making sushi, reading, podcasting, or trying something new.
  • Publications:

Tyler Shelby

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Epidemiology, public/global health, development of diagnostics for resource-limited settings, social determinants of health
  • Education:  Pittsburgh State University, B.S., 2016 (Biology, Chemistry) 
  • A little about me: I’ve lived the majority of my life in Southeast Kansas, so if you ever need a detailed description of a cornfield, I may be able to help you out. Outside of medical- and research-related activities, I’m a big fan of playing guitar and drums, traveling, bicycling, reading/writing, and eating food.
  • Publications:

Julio Silva

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak
  • Education:  University of California - Los Angeles, B.S., 2015 (Molecular, Cellular & Developmental Biology)
  • Publications:

Ronan Talty

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:
  • Education:  Northeastern University, B.S., 2017 (Behavioral Neuroscience) 
  • A little about me:
  • Publications:

Elizabeth Woo

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: I'm interested in the characterization and comparison of risk alleles in neurodegenerative diseases. Specifically, investigating the expression and function of those genes in women. Additionally, I would like to better understand the role of sex hormones as it relates to its potential protective capabilities.  
  • Education:  University of Chicago, B.A., 2017 (Economics) 
  • A little about me:  Born in Boston and grew up in Paris and the suburbs of Chicago, I love to learn foreign languages in my free time and to sing and jam out with friends. I love bad puns and will never turn down the opportunity to make one. My nicknames include, but are not limited to: Wootang, Liz-beth, Liz, Betty, and Woo.
  • Publications:

Sangwon Yun

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Immunobiology, immunoediting
  • Education:  Duke University, B.S., 2017 (Biology, Evolutionary Anthropology) 
  • A little about me: I was born in Seoul and raised in the San Francisco Bay Area. I continue to believe that I peaked in college when I performed in the Duke Jazz Ensemble with Gerald Clayton. My research interests focus on the intersection between immunology and oncology, particularly immune evasion. In my free time, I am usually reading, drinking coffee, beach bumming or struggling through yoga poses.
  • Publications:
Zekavat, Maryam

Maryam Zekavat

  • Year:  1
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Computational Biomedicine & Bioinformatics
  • Education:  Massachusetts Institute of Technology, B.S., 2015 (Biological Engineering) 
  • A little about me:  During my undergrad at MIT, I traveled to Roche in Basel, Switzerland and developed a mathematical model to explain the cholesterol dynamics of dry Age-related Macular Degeneration, a blinding and untreatable disease. After graduation, I worked as a computational biologist at the Broad Institute, using terabytes of whole-genome and whole-exome sequencing datasets to analyze both cholesterol and non-cholesterol pathways leading to cardiovascular disease. Besides medicine & research, I enjoy reading mystery books and I'm also an avid ice-skater (super excited about Yale's skating rink, aka "The Whale"). I'm also a 2017 Paul & Daisy Soros Fellow.
  • Publications:

Entering Class - 2016


Shawn Ahn

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Biomedical imaging and optics
  • Education:  U. of Illinois at Urbana-Champaign, B.S., 2016 (Electrical Engineering)
  • A little about me:  I was born and raised mostly in South Korea, but I lived in the west coast, midwest, and now I made it all the way to the east coast. I enjoy playing basketball whenever I have free time.

Emmanuella Asabor

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Health Policy and Management & History of Medicine; Refugee Health
  • Education:  Harvard College, B.A., 2013 (History and Science); University of Cambridge (M.Phil.), 2014, African Studies
  • A little about me:  I was born in London and spent my early childhood in Lagos, Nigeria, but I've called New York City home for most of my life. I enjoy traveling, cooking, ankara print, tea, Nigerian literature, and occasionally re-living my opera-singing past.

Ryan Chow

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Cancer heterogeneity, systems biology
  • Education:  Harvard College, B.A., 2016 (Human Developmental and Regenerative Biology)
  • A little about me:  During college, I minored in music! When I have the time, I also like submitting captions to the New Yorker Caption Contest.

Andrew Daniels

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Neuroscience, Immunology and Neuroimmunology
  • Education:  The Johns Hopkins University, B.A., 2014 (Neuroscience)
  • A little about me: I was born in Canada but have been in the United States since high school. While in college I was a member of the varsity track & field and cross country teams and in my free time I enjoy cooking, eating, running, and just about anything that involves being outdoors.

Luis Gonzalez

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  Genetics
  • Education:  University of California-L.A., B.S., 2015 (Biochemistry)
  • A little about me:

Michael Gormally

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Chemical Biology, Oncology, High-throughput Screening and Genomics.
  • Education:  Pomona College, B.A., 2011 (Chemistry); University of Cambridge, Ph.D., 2016 (Chemical Oncology)
  • A little about me:  I love the outdoors, particularly climbing up rocks, skiing down mountains and swimming in oceans. On a leisurely day I enjoy river punting with some cheese and champagne, and regularly partook in the Cambridge Punt Races down the River Cam.
Green, S

Sydney Rose Green

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest:  History of Medicine; Development of clinical intimate violence protocol
  • Education:  Harvard College, B.A., 2013 (History and Science)
  • A little about me:   A little about me: I am from Pembroke Pines, FL. I love knitting and watching college football (go Florida State!). Also, being from South Florida, I have soft spot in my heart for beaches, swampland, and cafecitos.

Kenneth Gunasekera

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Public Health; Infectious Disease; Pediatrics; Global Health
  • Education:  Yale University, B.S., 2015 (MB&B) 
  • A little about me:  I'm an MD-PhD student interested in studying the implementation of best practices in prevention, surveillance, and intervention to stop the spread of infectious disease. I have spent time doing work in Sri Lanka studying the social determinants of risk of HIV and traversing the tropical island paradise country. I love traveling, cooking, running, swimming, and spending time with friends and family (and also reading about all these things!


Grant Higerd

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Neuroscience, Immunology, Infectious Disease
  • Education:  University of California-L.A., B.S., 2015 (Neuroscience)
  • A little about me:  I grew up in Mammoth Lakes, CA where my family taught me to love ski racing, backpacking, and UCLA football. While at UCLA I was Captain of the rowing team. In my free time I enjoy doing anything that gets me outside, and traveling to the world's beautiful places.

Justin Johnson

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: 
  • Education:  Yale University, B.S., 2016 (MB&B)
  • A little about me:

Laurel Kaye

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Biomedical engineering and neuroscience
  • Education:  Duke University, B.S., 2015 (Physics)
  • A little about me:  I am crazy about the space program- I want to be an astronaut. I also love running, rowing and creative writing.

Sahana Kribakaran

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: 
  • Education:  University of California, L.A., B.S., 2015 (Neuroscience)
  • A little about me: 

Mytien Nguyen

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Host-microbe interactions; microbial pathogenesis in pulmonary diseases
  • Education:  Cornell University, B.S., 2014 (Biological Sciences, Microbiology)
  • A little about me: Born in Vietnam and raised in Vermont, I love family, food, and nature. My other loves are pandas, pugs, florals, and coloring books. My passions are social justice, environmental health justice, and education equity. Naturally, my apartment has an open door policy, especially to those who enjoy playing Exploding Kittens.

Samantha Olyha

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: stem cell biology, hematology/oncology 
  • Education:  Cornell University, B.S., 2014 (Biological Sciences)
  • A little about me:  I grew up in Poughkeepsie, NY and I have a fascination with turtles and tortoises! I also like to run, swim, and row, and am addicted to podcasts. After graduating, I spent 2 years as a grad student in the UK where I learned to fully appreciate tea, crumpets, and British English.  

Annsea Park

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: RNA biology, gene regulation 
  • Education:  University of California-Berkeley, B.A., 2014 (Molecular Cell Biology-Biochemistry)
  • A little about me: I was born in Ann Arbor, Michigan and raised in Seoul, South Korea. In high school, I went to India, Pakistan, and China where I became enthusiastic about volunteering. My hobbies include yoga, ballet, and playing card games (especially Exploding Kittens with my classmates!).

Eric Hoyeon Song

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Drug Delivery, Gene Delivery, Immunobiology
  • Education:  University of Southern California, B.S., 2014 (Biochemistry)
  • A little about me:

Alexandra Suberi

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Drug Delivery, Gene Delivery, Immunobiology
  • Education:  University of Maryland-College Park, B.S., 2016 (Biochemistry)
  • A little about me:  I like learning about permaculture and innovations in agriculture. I spend my free time hiking, cooking, and playing fetch with pups. 

Taylor Takasugi

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Drug Delivery, Gene Delivery, Immunobiology
  • Education:  University of Minnesota-Twin Cities, B.S., 2016 (Biochemistry, Genetics, Cell Biology)
  • A little about me:
Zhang, Ce

Ce Zhang

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: Drug Delivery, Gene Delivery, Immunobiology
  • Education: Pennsylvania State University, B.S., 2012 (Biology, Neuroscience Option)
  • A little about me:
Zogg, Cheryl

Cheryl Zogg

  • Year:  2
  • Mentor Information:  Barbara Kazmierczak 
  • Research interest: I am a health services researcher interested in the intersection of health policy and quality as it pertains to outcomes of surgical patients and differential access to care. From a methodological perspective, I am motivated by the adaptation and validation of novel statistical methodology with the overarching goal of improving approaches to large database research.
  • Education:  Macalester College, B.A., 2011 (ACS Honors Chemistry (Biochemistry) and Biology (Neurobiology); Johns Hopkins Bloomberg School of Public Health, M.S.P.H. and M.H.S., 2014 (Health Systems and Epidemiology).
  • A little about me:  I am a Midwesterner originally from Minneapolis/St. Paul, MN, who grew up as an avid ballet dancer. In my free time, I can usually be found curled up with a good book, traveling with friends, or enjoying the theater.
  • Publication:   Zogg CK, Jiang W, Ottesen TD, Shafi S, Schuster K, Becher B, Davis KA, Haider AH. Racial/Ethnic Disparities in Longer-term Outcomes Among Emergency General Surgery Patients: The Unique Experience of Universally-Insured Older Adults. Ann Surg 2017 [In Press].

Entering Class - 2015

Matthew Alsaloum

  • Year:  3
  • Mentor Information:  Stephen Waxman, INP
  • Education:  SUNY - Stony Brook, B.S., 2015 (Biology)

Jennifer Chen

  • Year:  3
  • Mentor Information:  Stephanie Eisenbarth, Immunobiology
  • Research interest: Immunobiology and molecular biology
  • Education:  Harvard University, B.A., 2015 (Human Development & Regenerative Biology)
  • A little about me:  I am part of the grad school a cappella group, The Citations!

Adriana Cherskov

  • Year:  3
  • Mentor Information:  Nenad Sestan, INP
  • Education:  Princeton University, B.A., 2014 (Molecular Biology)

Stefano Daniele

  • Year:  3
  • Mentor Information:  Nenad Sestan, INP
  • Research interest: Neuroscience, Neurodegeneration, Evolutionary Neurobiology, Glial Activation, Non-cell Autonomous Mechanisms, Protein Misfolding
  • Education:  Georgetown University, B.S., 2012 (Neurobiology)
  • A little about me: Soccer, Science, Salt

Tyrone DeSpenza

  • Year:  3
  • Mentor Information:  Kristopher Kahle, INP
  • Research interest:  I am interested in understanding the molecular genetics underlying neurodevelopmental disorders.
  • Education:  Dartmouth College, B.S., 2013 (Neuroscience)
  • A little about me:  I was born and raised in New Orleans, Louisiana. I love playing jazz music on my alto saxophone.

Matthew Dong

  • Year:  3
  • Mentor Information:  Sidi Chen, Genetics
  • Education:  U.C. Berkeley, B.S., 2012 (Molecular Biology)

Nicholas Economos

  • Year:  3
  • Mentor Information:  Peter Glazer, Genetics
  • Research interest: Immunology, DNA biology and repair
  • Education:  Tufts University, B.S., 2013 (Biology)
  • A little about me:  I'm from a small farm town in rural upstate New York and have since lived in Boston, London, and New York City before settling in New Haven. My hobbies include cooking, art/drawing, running, swimming, trampoline parks, and my pet iguana. On campus I have a tendency to join fringe medical interest groups including psychedelic medicine, wilderness medicine, and space medicine (which I co-founded during my first year).

Margret Erlendsdottir

  • Year:  3
  • Mentor Information:  Hongyu Zhao, Genetics
  • Education:  Yale University, B.S., 2015 (M.C.D.B.)

Carrie Flynn

  • Year:  3
  • Mentor Information:  Barbara Kazmierczak, Microbiology
  • Education:  Johns Hopkins, B.A., 2010 (Natural Sciences).  Rutgers University, M.S., 2015 (Biomedical Sciences)
  • Research interest:  Microbiology, antibiotic resistance, and the microbiome
  • A little about me:  I grew up in New Jersey and am interested in Infectious Disease. I love animals (especially horses and dogs), and am addicted to crossword puzzles and podcasts. I am also an activist by nature and am passionate about social justice issues.

James Garritano

  • Year:  3
  • Mentor Information:  Yuval Kluger, Pathology
  • Education:  U.C.L.A., B.S., 2011 (Biochemistry, Electrical Engineering)
  • A little about me:  I worked in UCLA's Center for Advanced Surgical and Interventional Technology (CASIT) for four years before coming to Yale. I helped design and characterize novel imaging systems, and developed mathematical models for surgical planning.

Casey Grun

  • Year:  3
  • Mentor Information:  Barbara Kazmierczak, Microbiology
  • Research interest: Novel antimicrobials, Microbial pathogenesis, Microbiome
  • Education:  Harvard University, B.S., 2014 (Biomedical Engineering)
  • A little about me:  I'm from the Pacific Northwest, so I love spending time outdoors—hiking, camping, and backpacking around New England. I also really like to cook—I specialize in baking funky chocolate chip cookies!

Danielle Miyagishima

  • Year:  3
  • Mentor Information:  Murat Gunel, Genetics
  • Research interest:  I'm interested in brain tumor genetics and small molecule targeting
  • Education:  New Mexico State University, B.S., 2015 (Biology)
  • A little about me:  I brought my three dogs (Laddie, Teddy, and Daisy) from New Mexico with me. I love the outdoors and am particularly fond of gardening, backpacking/hiking, swimming, running, and cycling. Fun fact: When I was 19, I got a Top Secret clearance from the US Government.

Raman Nelakanti

  • Year:  3
  • Mentor Information:  Andrew Xiao, Genetics
  • Education:  Stanford University, B.S., 2014 (M.C.D.B)
Park, Jonathan

Jonathan Park

  • Year:  3
  • Mentor Information:  Mark Gerstein, MB&B
  • Research interest:  Computational Biology
  • Education:   Yale University, B.S., 2014 (Molecular, Cellular and Developmental Biology)
  • A little about me: I was born in Chicago and grew up on Air Force bases in Japan and England.

Elias Quijano

  • Year:  3
  • Mentor Information:  Peter Glazer & Mark Saltzman, Genetics
  • Research interest:  Drug Delivery, Biomaterials, Gene Therapy, Gene Editing
  • Education:  Yale University, B.S., 2012 (B.M.E)
  • A little about me:  I was born in Newark, New Jersey, and raised in Hialeah, Florida. My hobbies include powerlifting, hiking, and reading. Before starting my MD/PhD, I spent some time doing research at Yale in drug and gene delivery under W. Mark Satlzman. 
  • Publications: Saucier-Sawyer J, Seo Y, Gaudin A, Quijano E, Song E, Sawyer A, Deng Y, Huttner A,  Saltzman WM. Distribution of polymer nanoparticles by convection-enhanced delivery to brain tumors. Journal of Controlled Release. 2016. PMID: 27063424.
  • Saucier-Sawyer J, Deng Y, Seo Y, Cheng C, Zhang J, Quijano E, Saltzman WM. Systemic Delivery of Blood-Brain Barrier Targeted Polymeric Nanoparticles Enhances Delivery to Brain Tissue. Journal of Drug Targeting. 2015. PMID: 26453169. PMCID: PMC4860350
  • McNeer NA, Anandalingam K, Fields RJ, Caputo C, Kopic S, Gupta A, Quijano E, Polikoff L, Kong Y, Bahal R, Geibel JP,  Glazer PM, Saltzman WM, Egan ME. Nanoparticles that deliver triplex-forming peptide nucleic acid molecules correct F508del CFTR in airway epithelium. Nature Communication. 2015. PMID: 25914116. PMCID: PMC4480796.
  • Fields RJ, Quijano E, McNeer NA, Caputo C, Bahal R, Anandalingam K, Egan ME, Glazer PM, Saltzman WM. Modified Poly(lactic-co-glycolic Acid) Nanoparticles for Enhanced Cellular Uptake and Gene Editing in the Lung. Advanced Healthcare Materials. 2014. PMID: 25156908.
  • Bahal R, Quijano E, McNeer NA, Liu Y, Bhunia DC, López-Giráldez F, Fields RJ, Saltzman WM, Ly DH, Glazer PM. Single-Stranded γPNAs for In Vivo Site-Specific Genome Editing via Watson-Crick Recognition. Current Gene Therapy. 2014. PMID: 25174576.
  • ields RJ, Cheng CJ, Quijano E, Weller C, Kristofik N, Duong N, Hoimes C, Egan ME, Saltzman WM. Surface modified poly(β amino ester)-containing nanoparticles for plasmid DNA delivery. Journal of Controlled Release. 2012. PMID: 23041278. PMCID: PMC3625648.

Diana Yanez

  • Year:  3
  • Mentor Information:  Oscar Colegio, Immunobiology
  • Education:  U.C.L.A, B.S., 2012 (M.C.D.B)

Entering class - 2014

Wardah Athar

  • Year:  4
  • Mentor Information:  Sreeganga Chandra, INP
  • Research interest:  Chandra lab
  • Education:  Georgetown University, B.S., 2013 (Neurobiology)
  • A little about me:  I grew up between Pakistan, Texas, Oman, and Saudi Arabia before settling in DC for college. I spent a year studying BME in Ireland and traveling around Europe post-graduation, after which I ended up in glamorous New Haven! In my spare time, I enjoy cooking, hiking, and spoiling my goofy rescue dog. 
  • Publication:  Kosmaczewski, S.G., Han, S.M., Han, B., Meyer, B., Baig, H.S., Athar, W., Lin_moore, A.T., Koelle, M.R., and Hammarlund, M. (2015). RNA ligation in neurons by RtcB inhibits axon regeneration. PNAS 11(27), 8451-6.

Daniel Barson

  • Year:  4
  • Mentor Information:  Michael Higley & Michael Crair, INP
  • Research interest:  Activity-dependent formation of neural circuity in development and disease and the role of glial cells in these processes.  Systems neuroscience techniques development.
  • Education: Princeton University, B.A., 2012 (Molecular Biology).  University of Cambridge, MPhil 2013 (Clinical Neurosciences)
  • A little about me:  I hail from the wilderness of the New York City suburbs, where I learned the survival skills necessary to take home the top prize (along with 3 other YSM2018ers (2 MudPhuds)) in the 1st Annual MedWar Massachusetts. When not bush-whacking or simulating emergent cricothyrotomies with fake blood made from ketchup, I fill my free time with cooking and indulging in New Haven's (surprisingly awesome) food scene, writing medicine-themed covers to hip-hop songs, adventuring throughout New England, and day-dreaming about the puppy that my partner and I will eventually adopt.  Preferred New Haven pizza:  Pepe's

Abigail Greene

  • Year:  4
  • Mentor Information:  Todd Constable, INP
  • Education:  Princeton University, B.A., 2012 (Psychology)

William Hancock-Cerutti

  • Year:  4
  • Mentor Information:  Pietro De Camilli, Cell Biology
  • Education:  Franklin & Marshall College, B.A., 2013 (Chemistry)

Corey Horien

  • Year:  4
  • Mentor Information:  Todd Constable & Laura Ment, INP
  • Research interest: Computational neuro, neuroimaging, developing brain (Constable and Ment labs)
  • Education:  Concordia College at Moorhead, B.A., 2013 (Biology)
  • A little about me:  Lover of all things Minnesota - go Vikings!

Woong Hwang

  • Year:  4
  • Mentor Information: Mustafa Khokha, Genetics
  • Education:  University of Rochester, B.S., 2013 (Neuroscience)

Ruoyi (Roy) Jiang

  • Year:  4
  • Mentor Information:  Steven Kleinstein, Pathology & Kevin O'Connor, Neurology
  • Education:  Yale University, B.S., 2014 (MCDB)

Daniel Kim

  • Year:  4
  • Mentor Information:  Akiko Iwasaki, Immunobiology
  • Research interest: Immunology
  • Education:  Harvard University, B.A., 2014 (Chemical & Physical Biology)
  • A little about me:  I love eating out and discovering new restaurants in New Haven!

Zachary Kloos

  • Year:  4
  • Mentor Information:  Christine Jacob-Wagner, Microbial Pathogenesis
  • Education:  Case Western Reserve University, B.S., 2011 (Biochemistry, Economics)
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Katherine Leiby

  • Year:  4
  • Mentor Information:  Laura Niklason, BME
  • Research interest: Lung tissue engineering (Laura Niklason’s lab)
  • Education:  Yale University, B.S., 2014 (BME)
  • A little about me: I grew up in a small town in Pennsylvania. In my free time, I love spending time outdoors, running, cooking, and playing the carillon (a set of giant bells hung in a tower). 
  • Publications:  Balestrini JL, Gard AL, Liu A, Leiby KL, Schwan J, Kunkemoeller B, Calle EA, Sivarapatna A, Lin T, Dimitrievska S, Campbell SG, Niklason LE. Production of decellularized porcine lung scaffolds for use in tissue engineering. Integr Biol (Camb). Dec 2015, 7(12):1598-610. PMID: 26426090. PMCID: PMC4666745. 
  • Calle EA, Mendez JJ, Ghaedi M, Leiby KL, Bove PF, Herzog EL, Sundaram S, and Niklason LE. Fate of distal lung epithelium cultured in a decellularized lung extracellular matrix. Tissue Eng Part A Original. May 2015, 21(11-12):1916-1928. PMID: 25789725. PMCID: PMC4449714.

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Jonathan Liang

  • Year:  4
  • Mentor Information:  Iain Fraser, Oxford; GPP-NIH
  • Education:  Yale University, B.S., 2013 (MB&B)
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Jonathan Marquez

  • Year:  4
  • Mentor Information:  Mustafa Khokha, Genetics
  • Research interest: Developmental and Cancer Biology (Khokha Lab)
  • Education:  Yale University, B.S., 2013 (MCDB)
  • A little about me: I am originally form Texas and enjoy soccer and bacon and I like turtles. 
  • Publications:  Pax7 is Regulated by cMyb During Early Neural Crest Development Through a Novel Enhancer. Stephanie Vadasz, Jonathan Marquez, Maria Tulloch, Natasha Shylo, and Martín I. García-Castro. 2013 Sept. Development 140: 3691-372.
  • SUMOylation of Pax7 is Essential for Neural Crest and Muscle Development. Zhidong Luan, Ying Liu, Timothy J. Stuhlmiller, Jonathan Marquez, and Martín I. García-Castro. 2013 May. Cellular and Molecular Life Sciences 70: 1793-1806.
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Sarah Meller

  • Year:  4
  • Mentor Information: Charles Greer, INP
  • Education:  Carlton College, B.A., 2013 (Biology)
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Jessica Minor

  • Year:  4
  • Mentor Information:  Richard Bribiescas, Anthropology
  • Research interest: Biocultural Anthropology
  • Education:  Washington University, B.A., 2014 (Spanish & International Studies)
  • A little about me: With insights from the biomedical and social sciences, I explore the health consequences of social adversity in immigrant communities through the lens of life history theory. Outside of research, I volunteer as a Behavioral Health Psychoeducator at HAVEN and serve on the medical school's Committee for Diversity, Inclusion, and Social Justice. I also co-founded www.SystemicDisease.com, an online community addressing unconscious bias in medicine.
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Alyssa Mitson-Salazar

  • Year:  4
  • Mentor Information:  Ruslan Medzhitov, Immunobiology
  • Research interest: Allergic host defenses (Medzhitov lab)
  • Education:  Yale University, B.S., 2013 (Biology)
  • A little about me:  Originally from rural Colorado, I fell in love with the type 2 immune system and the mechanisms of allergic disease during my time as a postbac at the NIH. When I'm not in the hospital or in lab, I spend my free time hiking the state parks of Connecticut, eating tacos, and playing with my 3-legged, 1-eared cat.
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Micha Sam Brickman Raredon

  • Year:  4
  • Mentor Information:  Laura Niklason, BME
  • Education:  Yale University, B.S., 2013 (History of Art, BME), Massachusetts Institute of Technology, 2014, Materials Science and Engineering
  • A little about me:   I play the cello and I try to go backpacking when possible.
  • Publications:   
  • “Tissue-engineered lungs for in vivo implantation.” Science, 2010. 
  • “A Call to Craft.” Science Translational Medicine, 2014 
  • “Alveolar epithelial differentiation of human induced pluripotent stem cells in a rotating bioreactor.” Biomaterials, 2014. 
  • “A rotating bioreactor for scalable culture and differentiation of respiratory epithelium.” Cell Medicine, 2015.
  • "Biomimetic Culture Reactor for Whole-Lung Engineering." BioResearch Open Access, 2016.
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Bingchan (Catherine) Xie

  • Year:  4
  • Mentor Information:  Jordan Pober, Immunobiology
  • Education:  California Institute of Technology, B.S., 2013 (Bioengineering & English); University of Cambridge, Master of Philosophy with distinction (Clinical Science), 2014 
  • A little about me: Growing up and living in four different countries (China, Australia, Canada, and the USA), I developed a passion for scientific discovery to better the human condition in addition to gaining a greater understanding of and appreciation for diversity and culture. As an MD-PhD at Yale, I am able to build the ideal foundation to become a physician-scientist, bridging the boundaries of the lab and clinic. In my spare time, I enjoy exploring the outdoors and expanding my culinary skills. 
  • Publication: Xie, C. B. et al. Regulation of aldosterone secretion by Ca V 1.3. Sci. Rep. 6, 24697; doi: 10.1038/srep24697 (2016).

Entering class - 2013

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Shivani Bhatt

  • Year:  5
  • Graduate Program:  INP
  • Mentor Information: Kelly Cosgrove, Psychiatry
  • Research interest: PET neuroimaging of PTSD and addiction
  • Education:  Yale University, B.S., 2013 (Biology)
  • A little about me: I grew up in the suburbs of Minneapolis but I LOVE New Haven and having been here for 7 years, I consider it a second home. Late-night eats? Best study spots? Community outreach opportunities? Ask me and I'm happy to give you more information than you ever wanted. At the medical school I am involved with Yale Medical Student Psychiatry Association, Refugee Health Clinic/Wednesday Evening Clinic, US Health Justice Collaborative, Committee on Women and Minority Support, and the Ultrasounds! a cappella group for health professions students. Off campus you can find me singing, reading news and long-form articles, soaking up the sun, and hanging out with the lovely friends I've met in my time here. 
  • Publications: Dhande OS, Bhatt S, Anishchenko A, Elstrott J, Iwasato T, Swindell EC, Xu HP, Jamrich M, Itohara S, Feller MB, Crair MC: Role of adenylate cyclase 1 in retinofugal map development. J Comp Neurol. 2012 May 1.
  • Dhande OS, Hua EW, Guh E, Yeh J, Bhatt S, Zhang Y, Ruthazer ES, Feller MB, Crair MC: Development of single retinofugal axon arbors in normal and β2 knock-out mice. J Neurosci. 2011 Mar 2.
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Swethasri Dravida

  • Year:  5
  • Graduate Program:  INP
  • Mentor Information:  Joy Hirsch, Psychiatry
  • Education:  MIT, B.S., 2013 (Brain & Cognitive Sciences, Mathematics)
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Erin Feeney

  • Year:  5
  • Graduate Program:  INP
  • Mentor Information: Philip Corlett, Psychiatry 
  • Education:  Davidson College, B.S., 2011 (Neuroscience)
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Amanda Jeng

  • Year:  5
  • Graduate Program: INP
  • Mentor Information:  Anthony Koleske, MB&B
  • Education:  Washington University, B.A., 2010 (Biology)
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Alanna Kaplan

  • Year:  5
  • Graduate Program: Experimental Pathology
  • Mentor Information:  Peter Glazer, Genetics 
  • Research interest: DNA repair inhibitors for cancer therapy
  • Education:  Dartmouth College, B.A., 2011 (Neuroscience)
  • A little about me:  I grew up in Chappaqua, NY and spent a few years in Washington, DC before coming to Yale. At Yale, I volunteer at HAVEN free clinic. I also enjoy yoga, running, baking, and checking out the New Haven restaurant scene. Feel free to contact me (alanna.kaplan@yale.edu) with any questions!
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Irina Krykbaeva

  • Year:  5
  • Graduate Program: Experimental Pathology
  • Mentor Information:  Marcus Bosenberg, Pathology
  • Education:  Brandeis University, B.S., 2011 (Neuroscience)
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Don Tianmu Li

  • Year:  5
  • Graduate Program: Cell Biology
  • Mentor Information:  Jonathan Bogan, Cell Biology 
  • Research interest: Molecular mechanisms of GLUT4 translocation in diabetes
  • Education:  Yale University, B.S., 2013 (Biochemistry)
  • A little about me : I am interested in the field of pediatric orthopedics clinically and in the disciplines of cell biology and biochemistry scientifically. Otherwise I enjoy teaching and mentoring and have been heavily involved in both undergraduate pre-med advising and the Students Helping Students program in the medical school. Less productively, I also like to hang out and play video games. I am more than happy to talk about my experiences in the program or about any other topic that you might be interested in! 
  • Publications:  Belman JP, Bian RR, Habtemichael EN, Li DT, Jurczak MJ, Alcázar-Román A, McNally LJ, Shulman GI, Bogan JS. Acetylation of TUG protein promotes the accumulation of GLUT4 glucose transporters in an insulin-responsive intracellular compartment. J Biol Chem. 2015 Feb 13;290(7):4447-63. doi: 10.1074/jbc.M114.603977. Epub 2015 Jan 5. PubMed PMID: 25561724; PubMed Central PMCID: PMC4326849.
  • Parker CG, Dahlgren MK, Tao RN, Li DT, Douglass EF Jr, Shoda T, Jawanda N, Spasov KA, Lee S, Zhou N, Domaoal RA, Sutton RE, Anderson KS, Jorgensen WL, Krystal M, Spiegel DA. Illuminating HIV gp120-Ligand Recognition through Computationally-Driven Optimization of Antibody-Recruiting Molecules. Chem Sci. 2014 Jun 1;5(6):2311-2317. PubMed PMID: 25379167; PubMed Central PMCID: PMC4217211.
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George Linderman

  • Year:  5
  • Graduate Program:  Applied Mathmatics
  • Mentor Information:  Yuval Kluger, Pathology and Ronald Coifman, Mathematics 
  • Research interest: Techniques for dimensionality reduction of biomedical data
  • Education:  Case Western Reserve University, B.S., 2013 (Biomedical Engineering)
  • A little about me: I was born in Cleveland, and when I was eight, my family and I moved to Albania where I lived until college. It was there that I began to program and was exposed to medicine through my father, a general surgeon. I derive great joy from learning, particularly in the fields of mathematics, medicine, Chinese, and aikido. Otherwise, I love to spend time with the wonderful people I am blessed to call my friends (including the people on this very page!).
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William Meyerson

  • Year:  5
  • Graduate Program: CB&B
  • Mentor Information:  Mark Gerstein, MB&B 
  • Research interest: Methods development in computational genomics
  • Education:  Yale University, B.S., 2013 (MB&B)
  • A little about me:  I love to learn about anything and explore ideas from multiple perpsectives. I enjoy conversations, reading, running, oratory, and games involving creative problem-solving. Washington, D.C. was my home growing up and have found another here at Yale. If I can find the wisdom that I seek, I hope to someday participate in global conversations about the choices that we as societies and as individuals face that impact health and well-being. Feel free to email me at william.meyerson@yale.edu 
  • Publication: Wang, D., Yan, K.K., Sisu, C., Cheng, C., Rozowsky, J., Meyerson, W., and Gerstein, M.B. (2015) Loregic: a method to characterize the cooperative logic of regulatory factors. PLoS Comput. Biol. 11, e1004132.
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Minh Than

  • Year:  5
  • Graduate Program:  Genetics
  • Mentor Information:  Richard Lifton, Genetics
  • Education:  University of Colorado at Boulder, B.A., 2012 (Molecular Biology)
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John Walsh

  • Year:  5
  • Graduate Program: BME
  • Mentor Information:  Fahmeed Hyder, BME
  • Education:  Florida State University, B.S., 2010 (Chemical Engineering)

Entering Class-2012

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Gregory Breuer

  • Year:  6
  • Graduate Program: Experimental Pathology
  • Mentor Information: Ranjit Bindra, Pathology
  • Education:  University of Connecticut, B.S., 2012 (Biophysics and Computer Science)
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Nashid Chaudhury

  • Year:  6
  • Graduate Program: INP
  • Mentor Information: Susumu Tomita, Physiology
  • Research interest:  Synaptic transmission and plasticity
  • Education: Barnard College, B.A., 2011 (Neuroscience & Behavior)
  • A little about me:  My family is originally from Bangladesh, and I was born & raised in Queens, NYC. 
  • Publication: Schobel S.A., Chaudhury N.H., Khan U.A., Paniagua B., Styner M.A., Asllani I., Inbar B.P., Corcoran, C.M., Lieberman J.A., Moore H., Small S.A. (2013) Imaging Patients with Psychosis and a Mouse Model Establishes a Spreading Pattern of Hippocampal Dysfunction and Implicates Glutamate as a Driver. Neuron. 78:81-93. PMID:  23583108
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William Culligan

  • Year:  6
  • Graduate Program: MB&B
  • Mentor Information: Matthew Simon, MB&B 
  • Research interest: Chemical biology of chromatin and lncRNAs
  • Education: University of Notre Dame, B.S., 2010 (Chemistry)
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Jessica Johnston

  • Year:  6
  • Graduate Program: Cell Biology
  • Mentor Information: Megan King, Cell Biology 
  • Research interest:  My current work in Megan King's lab (Cell Biology) involves biophysical approaches (force spectroscopy and quantitative fluorescence microscopy) to characterize the fundamental components of nuclear mechanics. I am particularly interested in how nuclear mechanics is altered in cancer metastasis, and how these altered mechanical states might influence DNA damage/repair.
  • Education: University of Utah, B.S., 2012 (Physics) 
  • A little about me:  A native Californian (but truly a Utahn at heart), I enjoy any opportunity to be outside, preferably hiking, backpacking, biking, rockclimbing, or skiing in the mountains. If I am not in lab or engaging in one of these activities, you will likely find me a) with friends, b) reading a book, or c) napping.
  • Publication:  Waite AJ, Frankel NW, Dufour YS, Johnston JF, and Emonet T. 2016. Non-genetic diversity modulates population performance. Molecular Systems Biology. In revision. 
  • Schniederberend M, Johnston JF, Emonet T, and Kazmierczak BI. 2016. In preparation.
  • Johnston JF, King MC. 2016. Thar she blows! Causes and consequences of nuclear rupture. Review. Cell. Submitted.

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Angela Lee

  • Year:  6
  • Graduate Program: INP
  • Mentor Information: Marina Picciotto, INP
  • Education: University of Maryland College Park, B.S., 2011 (Physiology, Neuroscience and Psychology)

Young Lim

  • Year:  6
  • Graduate Program: Genetics
  • Mentor Information: Keith Choate, Genetics, Pathology
  • Education: Yale University, B.S., 2011 (Biophysics)

Christina Lin

  • Year:  6
  • Graduate Program:  Microbiology
  • Mentor Information: Barbara Kazmierczak, Microbial Pathogenesis 
  • Research interest:  I am studying the regulation and expression of the Type III Secretion System in Pseudomonas aeruginosa.
  • Education: Yale University, B.S., 2011 (MB&B)
  • A little about me:  I'm interested in global health and infectious diseases. I love to create art, try out new cuisines, play video/board games. I'm also trying to slowly improve my half-marathon times.

Jacob Lister

  • Year:  6
  • Graduate Program: INP
  • Mentor Information:  Michael Crair, Neuroscience
  • Education: Middlebury College, B.A., 2011 (Philosophy)
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Alice Lu

  • Year:  6
  • Graduate Program:  Genetics
  • Mentor Information:  Natalia Ivanova, Genetics 
  • Research interest:  Stem cells
  • Education: Cornell University, B.S., 2010 (Biology) 
  • Publications:  Idevall-Hagren, O., Lu, A., Xie, B., & De Camilli, P. (2015). Triggered Ca2+ influx is required for extended synaptotagmin 1-induced ER-plasma membrane tethering. The EMBO Journal, 34(17), 2291–2305. http://doi.org/10.15252/embj.201591565
  • Courchaine, E. M., Lu, A., & Neugebauer, K. M. (2016). Droplet organelles? The EMBO Journal, e201593517–10. http://doi.org/10.15252/embj.201593517
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Matthew Meizlish

  • Year:  6
  • Graduate Program: Immunobiology
  • Mentor Information: Ruslan Medzhitov, Immunobiology
  • Education: Yale University, B.A., 2011 (History)
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Kevin Perkins

  • Year:  6
  • Graduate Program: MCDB
  • Mentor Information: Ronald Breaker, MCDB
  • Education: Rice University, B.S., 2012 (Biomedical Engineering)
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Lorenzo Sewanan

  • Year:  6
  • Graduate Program: BME
  • Mentor Information: Stuart Campbell, BME 
  • Research interest: Cardiac biomechanics and tissue engineering
  • Education: Trinity College, B.S., 2012 (Engineering and Physics) 
  • A little about me:  Born and raised in Suriname, adolescent years in Queens, learning to love New England one day a time.
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Durga Thakral

  • Year:  6
  • Graduate Program: Genetics
  • Mentor Information: Marcus Bosenberg, Pathology
  • Research interest:  Human Genetics
  • Education: Yale University, B.S., 2012 (Molecular Biophysics and Biochemistry) 
  • A little about me:  I study the genetics of congenital diseases using computational methods. 
  • Publication: Hakim, A., J.B. Nguyen, K. Basu, D.F. Zhu, D. Thakral, P.L. Davies, F.J. Isaacs, Y. Modis, and W. Meng. 2013. Crystal Structure of an insect antifreeze protein and its implications for ice binding. J Biol Chem. 288:12295-304. PMID: 23486477
  • Nkansah MK, Thakral D, Shapiro EM. (2011). Magnetic poly(lactide-co-glycolide) and cellulose particles for MRI-based cell tracking. Magnetic Resonance in Medicine. (65)(6): 1776-85.
    Hakim A, Nguyen JB, Basu K, Zhu DF, Thakral D, Davies PL, Isaacs FJ, Modis Y, Meng W. (2013). Crystal Structure of an insect antifreeze protein and its implications for ice binding. Journal of Biological Chemistry. (288)(17):12295-304.
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Andrew Timberlake

  • Year:  6
  • Graduate Program: Genetics
  • Mentor Information:  Richard Lifton, Genetics
  • Research interest:  I'm beginning my second year in the Lifton Lab, studying the genetics of craniofacial birth defects using next generation sequencing technologies.
  • Education: Yale University, B.S., 2011 (Molecular Biophysics and Biochemistry) 
  • A little about me: Originally from Mountain Brook, Alabama, I've now spent 9 years in New Haven. I love to run, play the drums, and spend time in New York City. I will be applying to Plastic Surgery programs after completing my PhD. 
  • Publication:   Chin A, Svejda B, Gustaffson BI, Granlund AB, Sandvik AK, Timberlake A, Sumpio B, Pragner R, Modlin IM, Kidd M. (2012) The role of mechanical forces and adenosine in the regulation of intestinal enterochromaffin cell serotonin secretion. Am J Physiol Gastrointest Liver Physiol: 302(3):G397-405. PMID: 22038827
  • Svejda, B.*, M. Kidd*, A. Timberlake, K. Harry, A. Kazberouk, S. Schimmack, B. Lawrence, R. Pfragner, and I.M. Modlin. 2013. Serotonin and the 5-HT7 receptor: The link between hepatocytes, IGF-1 and small intestinal neuroendocrine tumors. Cancer Sci. Epub ahead of print. PMID: 23578138. 
  • Flowers GP, Timberlake AT, McLean KC, Monaghan JR, Crews CM. (2014) Highly efficient targeted mutagenesis in axolotl using Cas9 RNA-guided nuclease. Development: 141(10):2165-71. PMID: 24764077
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Rebecca Treger

  • Year:  6
  • Graduate Program: Immunobiology
  • Mentor Information:  Akiko Iwasaki, Immunobiology
  • Education: Yale University, B.S., 2012 (Biology) 
  • Publications: Treger, Rebecca  S, Cook AG, Rai G, Maloney DJ, Simeonov A, Jadhav A, Thomas CJ, Williams DL, Cappello M, Vermeire JJ (2012). Oxadiazole 2-oxides are toxic to the human hookworm, Ancylostoma ceylanicum, however glutathione reductase is not the primary target. International Journal of Parasitology: Drugs and Drug Resistance, 2: 171-177. PMID 22844653
  • Treger, Rebecca S, Otchere J, Keil MF, Quagraine JE, Rai G, Mott BT, Humphries, DL, Wilson M, Cappello M, Vermeire JJ (2013). Short Report: In vitro Screening of Compounds against Laboratory and Field Isolates of Human Hookworm Reveals Quantitative Differences in Anthelminthic Susceptibility. American Journal of Tropical Medicine and Hygiene
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Jessica Ye

  • Year:  6
  • Graduate Program: Immunobiology
  • Mentor Information:  Ruslan Medzhitov, Immunobiology
  • Education: Cornell University, B.A., 2012 (Biology and Chemistry)
Ying new

Lee Ying

  • Year:  6
  • Graduate Program: Pharmacology
  • Mentor Information:  John Hwa & Kathleen Martin
  • Education: University of California, Berkeley, B.A., 2011 (Molecular and Cellular Biology)
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Laura Yockey

  • Year:  6
  • Graduate Program: Immunobiology
  • Mentor Information:  Akiko Iwasaki, Immunobiology
  • Education: Washington University, B.A., 2011 (Biology) 
  • Publication: Yockey, Laura J, Demehri S, Turkoz M, Turkoz A, Ahern PP, Jassim O, Manivasagam S, Kearney JF, Gordon JI, Kopan R. (2013) The Absence of A Microbiota Enhances Tslp Expression in Mice With Defective Skin Barrier But Does Not Affect The Severity Of Their Allergic Inflammation. J Invest Dermatol. Epub ahead of print. PMID: 23698100
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Mark Youngblood

  • Year:  6
  • Graduate Program: INP
  • Mentor Information: Murat Gunel, Neurobiology
  • Research interest:  Genomic drivers of meningioma and other brain tumors
  • Education:  Georgia Tech, B.S., 2007 (Electrical Engineering) 
  • A little about me:  My wife and I are originally from the south-east and are currently living in Hamden. We recently had a son named Sebastian. We're interested in camping and hiking in New England and other areas. 
  • Publications:  Clark VE, Serin Harmanci A, Bai H, Youngblood MW, Lee T, Baranoski J, Ercan-Sencicek AG, Abraham BJ, Weintraub A, Hnisz D, Simon M, Krischek B, Günel JM, Erson-Omay EZ, Henegariu O, Carrión-Grant G, Mishra-Gorur K, Goldmann JE, Schramm J, Vortmeyer AO, Piepmeier JM, Bilgüvar K, Yasuno K, Young RA, Günel M. “Recurrent, somatic mutations in POLR2A define a distinct subset of meningiomas.” Nature Genetics. In Press.
  • Youngblood MW, Chen WC, Mishra AM, Enamandram S, Sanganahalli BG, Motelow JE, Bai HX, Frohlich F, Gribizis A, Lighten A, Hyder F, Blumenfeld H. (2015) “Rhythmic 3-4 Hz discharge is insufficient to produce cortical BOLD decreases in generalized seizures.” Neuroimage, 109:368–377.
  • Yoo JY, Farooque P, Chen W, Youngblood MW, Zaveri HP, Gerrard JL, Spencer DD, Hirsch LJ, Blumenfeld H. (2014) “Ictal spread of medial temporal lobe seizures with and without secondary generalization: an intracranial EEG analysis.” Epilepsia, 55(2):289-295
  • Lee HW, Youngblood MW, Farooque P, Han X, Jhun S, Goncharova I, Vives K, Spencer DD, Zaveri H, Hirsch LJ, Blumenfeld H. (2013) “Seizure localization using three-dimensional surface projections of intracranial EEG power.” Neuroimage, 83:616-626.
  • Youngblood MW, Han X, Farooque P, Jhun S, Bai X, Yoo JY, Blumenfeld H. (2013) “Intracranial EEG Surface Renderings: New Insights into Normal and Abnormal Brain Function.” The Neuroscientist, 19(3), 238-247.

Entering Class - 2011

Alexandra Albert

  • Year:  7
  • Graduate Program: Cell Biology
  • Mentor Information: Don Nguyen, Pathology 
  • Research interest: Adaptive responses to cellular stress in lung adenocarcinoma progression and metastasis (Nguyen Lab)
  • Education: University of Maine, B.S., 2009 (Biochemistry, Mol. Cel. Biology)
  • A little about me: I'm a born and raised Mainer with a interest in understanding how cancer cells adapt to new environments. 

Dipankan Bhattacharya

  • Year: 7
  • Graduate Program: Cell Biology
  • Mentor Information: Mustafa Khokha, Genetics 
  • Research Interest: Developmental biology and understanding the molecular and cellular function of genes implicated in human disease
  • Education: UC Berkeley, B.A., 2010 (Molecular Cellular Biology - Genetics, Genomics & Development)
  • A little about me: I am from the Bay Area, California. I enjoy meeting people and often give hugs in lieu of handshakes. I love martial arts, making parody videos, anything active like hiking, rock climbing, paintballing, skiing, etc. and of course... GO BEARS!!! 
  • Publications: Jonas S, Zhou E, Deniz E, Huang B, Chandrasekera K, Bhattacharya D, Wu Y, Fan R, Deserno TM, Khokha MK, Choma MA, 2013, A novel approach to quantifying ciliary physiology: microfluidic mixing driven by a ciliated biological surface. Lab on a chip, 13(21):4160-3. PMID: 23970350; PMCID: PMC3856250 
  • del Viso F, Bhattacharya D, Kong Y, Gilchrist MJ, Khokha MK, 2012, Exon capture and bulk segregant analysis: rapid discovery of causative mutations using high-throughput sequencing. BMC genomics, 13:649. PMID: 23171430; PMCID: PMC3526394 
  • Jonas S, Bhattacharya D, Khokha MK, Choma MA, 2011, Microfluidic characterization of cilia-driven fluid flow using optical coherence tomography-based particle tracking velocimetry. Biomedical optics express, 2(7):2022-34. PMID: 21750777; PMCID: PMC3130586

Sean Bickerton

  • Year: 7
  • Graduate Program: BME
  • Mentor Information: Tarek Fahmy, BME
  • Education: University of Amherst, B.S., 2010 (Biochemistry, Chemistry)

Dimitri de Kouchkovsky

  • Year: 7
  • Graduate Program: Immunobiology
  • Mentor Information: Carla Rothlin, Immunobiology 
  • Research interest:  I am interested in how the immune system is controlled. Specifically I am interested in how different types of immune response (e.g. anti-bacterial or anti-parasitic) are chosen and what factors control this as well as how immune tolerance (the ability of the body to not attack self tissues) is maintained.
  • Education: UC Berkeley, B.A., 2010 (Mol. Cel. Biology)
  • A little about me:  I was born in France and moved to the Bay Area when I was 11. While at UC Berkeley I started working in a lab at UCSF. In this lab I was first exposed to immunology and quickly became fascinated by this field and its position at the intersection of science and medicine. I decided to pursue a MD-PhD to be able to continue to study the immune system and understand when its dysfuntion causes disease. When not in lab I like to go on long bike rides around New Haven, play rugby and cook.  
  • Publication:  Chan PY, Carrera Silva EA, De Kouchkovsky D, Joannas LD, Hao L, Hu D, Huntsman S, Eng C, Licona-Limón P, Weinstein JS, Herbert DR, Craft JE, Flavell RA, Repetto S, Correale J, Burchard EG, Torgerson DG, Ghosh S, Rothlin CV. The TAM family receptor tyrosine kinase TYRO3 is a negative regulator of type 2 immunity. Science. 2016 Apr 1;352(6281):99-103.

Ramak Khosravi

  • Year: 7
  • Graduate Program: BME
  • Mentor Information: Jay Humphrey, BME 
  • Research interest: Computational model-driven design of tissue engineered vascular grafts
  • Education: University of Toronto, B.S., 2011 (Developmental Biology Specialist)
  • A little about me:  I'm originally from Toronto, Canada and chose the MD-PhD program because I am interested in applying what we learn from developmental biology and organogenesis to regenerative medicine. My PhD focuses on understanding how the biomechanical properties of readily implantable polymeric scaffolds influence long-term graft growth and remodeling. This is a collaborative project spanning across biomedical engineering, cardiovascular surgery, and biomaterial design. I hope to translate this research into improved outcomes in congenital heart surgery in my future career. 
  • Publication : Figley SA, Khosravi R, Legasto JM, Tseng YF, Fehlings MG, 2014, Characterization of Vascular Disruption and Blood-Spinal Cord Barrier Permeability following Traumatic Spinal Cord Injury. J Neurotrauma, 31(6):541-52. PMID: 24237182; PMCID: PMC3949504
  • Udelsman, B.V., Khosravi, R., Miller, K.S., Dean, E.W., Bersi, M.R., Rocco, K., Yi, T., Humphrey, J.D., and Breuer, C.K. (2014) Characterization of evolving biomechanical properties of tissue-engineered vascular grafts in the arterial circulation. Journal of Biomechanics 47(9):2070-9.
  • Kurobe, H., Tara, S., Maxfield, M.W., Rocco, K.A., Bagi, P.S., Yi, T., Udelsman, B.V., Dean, E.W., Khosravi, R., Powell, H.M., Shinoka, T., Breuer, C.K. (2015) Comparison of the biological equivalence of two methods for isolating bone marrow mononuclear cells for fabricating tissue-engineered vascular grafts. Tissue Engineering Part C 21(6):597-604.
  • Miller, K.S., Khosravi, R., Breuer, C.K., Humphrey, J.D. (2015) A hypothesis-driven parametric study of effects of polymeric scaffold properties on tissue engineered neovessel formation. Acta Biomaterialia 11:283-94.
  • Khosravi, R., Miller, K.S., Best, C.A., Shih, Y.C., Lee, Y.U., Yi, T., Shinoka, T., Breuer, C.K., Humphrey, J.D. (2015) Biomechanical diversity despite mechano- biological stability in tissue engineered vascular grafts two years post- implantation. Tissue Engineering Part A 21(9-10):1529-38.
  • Khosravi, R., Best, C.A., Allen, R.A., Stowell, C.E.T., Onwuka, E., Zhuang, J.J., Lee, Y.U., Yi, T., Bersi, M.R., Shinoka, T., Humphrey, J.D., Wang, Y., Breuer, C.K. (2016) Long-term functional efficacy of a novel electrospun poly(glycerol sebacate)-based vascular graft in mice. Annals of Biomedical Engineering (Epub ahead of print).

Amanda King

  • Year: 7
  • Graduate Program: BME
  • Mentor Information: Mark Saltzman, BME
  • Education: University of Pennsylvania, B.S., 2011 (Chemical Engineering)

Ashton Lai

  • Year: 7
  • Graduate Program: MCDB 
  • Research interest: Chemical Biology, Novel Therapeutics
  • Mentor Information: Craig Crews, MCDB
  • Education: Duke University, B.S., 2011 (Chemistry)
  • Publication:  Lai AC, Toure M, Hellerschmied D, Salami J, Jamie-Figueroa S, Ko E, Hines J, Crews CM. Modular PROTAC Design for the Degradation of Oncogenic BCR-ABL. Angew Chem Int Ed Engl, 2016. 55(2): p. 807-10. PMID: 26593377; PMCID: PMC4733637

Jonathan Levinsohn

  • Year: 7
  • Graduate Program: Genetics
  • Mentor Information: Keith Choate, Pathology 
  • Research Interest: Choate Lab; human genetics, mosaic disease.
  • Education: Williams College, B.A., 2010 (Biology and Chemistry)
  • A little about me:  I enjoy playing soccer and running.
  • Publications:  Levinsohn JL, Sugarman JL, Yale Center for Mendelian Genomics, McNiff MJ, Antaya RJ, Choate KA (2016) Somatic Mutations in NEK9 Cause Nevus Comedonicus. Am J Hum Genet. 2016 May 5. 
  • Walker S, Wang C, Walradt T, Hong BS, Tanner JR, Levinsohn JL, Goh G, Subtil A, Lessin SR, Heymann WR, Vonderheid EC, King BA, Lifton RP, Choi J (2015) Identification of a gain-of-function STAT3 mutation (p.Y640F) in lymphocytic variant hypereosinophilic syndrome. Blood. 2015 Dec 23
  • Goh G, Walradt T, Markarov V, Blom A, Riaz N, Doumani R, Stafstrom K, Moshiri A, Yelistratova L, Levinsohn J, Chan TA, Nghiem P, Lifton RP, Choi J (2015) Mutational landscape of MCPyV-positive and MCPyV-negative merkel cell carcinomas with implications for immunotherapy. Oncotarget. 2015 Dec 7.
  • Levinsohn JL, Sugarman JL, Bilguvar K, McNiff JM, Yale Center for Mendelian Genomics, Choate KA (2015) Somatic V600E BRAF mutation in linear and sporadic syringocystadenoma papilliferum. Journal for Invest Dermatol. 2015 Oct.
  • Levinsohn JL, McNiff JM, Antaya RJ, Choate KA (2015) A Somatic p.G45E GJB2 Mutation Causing Porokeratotic Eccrine Ostial and Dermal Duct Nevus. JAMA Dermatol. 2015 Jun.
  • Levinsohn JL, Teng J, Craiglow BG, Loring EC, Burrow TA, Mane SS, Overton JD, Lifton RP, McNiff JM, Lucky AW, Choate KA (2013) Somatic HRAS p.G12S Mutation Causes Woolly Hair and Epidermal Nevi. Journal for Invest Dermatol. 2014 Apr.
  • Levinsohn JL, Tian LC, Boyden LM, McNiff JM, Narayan D, Loring ES, Yun D, Sugarman JL, Overton JD, Mane SM, Lifton RP, Paller AS, Wagner AM, Antaya RJ, Choate KA (2013) Whole exome sequencing reveals somatic mutations in HRAS and KRAS which cause nevus sebaceus. Journal for Invest Dermatol. 133:827-30.

Alice Li

  • Year: 7
  • Graduate Program: Cell Biology
  • Mentor Information: Daniel Colon-Ramos, Cell Biology
  • Education: UCLA, B.S., 2011 (Physiological Sciences)
  • Publications: Reyes-Lamothe R, Tran T, Meas D, Lee L, Li AM, Sherratt DJ, Tolmasky ME, 2014, High-copy bacterial plasmids diffuse in the nucleoid-free space, replicate stochastically and are randomly partitioned at cell division. Nucleic acids research, 42(2):1042-51. PMID: 24137005; PMCID: PMC3902917 
  • Mineur, Y.S., A. Obayemi, M.B. Wigestrand, G.M. Fote, C.A. Calarco, A.M. Li, M.R. Picciotto. (2013) Cholinergic signaling in the hippocampus regulates social stress resilience and anxiety- and depression-like behavior. PNAS. 110(9):3573-3578. PMID: 23401542. 
  • Rajic, Z., Tovmasyan, A., Spasojevic, I., Sheng, H., Lu, M., Li, A.M., Gralla, E.B., Warner, D.S., Benov, L., Batinic-Haberle, I. (2012) A new SOD mimic, Mn(III) ortho N-butoxyethylpyridylporphyrin, combines superb potency and lipophilicity with low toxicity. Free Radic. Biol. Med. 52:1828-1834. PMID: 22336516.

Kavita Mistry

  • Year: 7
  • Graduate Program: C&M Physiology
  • Mentor Information: Michael Caplan, C&M Physiology 
  • Research interest: My project seeks to understand the molecular basis for autosomal dominant polycystic kidney disease.
  • Education: Yale, B.A., 2011 (English Literature, MB&B)
  • A little about me: My activities in the medical school include organizing our MSTP clinical reasoning workshop series and co-directing the YSM Peer Tutoring Program. I'm also very involved with the undergraduates -- in addition to teaching a section for the undergraduate physiology course, I serve as a Berkeley College graduate affiliate and I dance on an undergraduate Indian folk dance team. My informal interests include gardening, playing the piano, horseback riding, and listening to audiobooks.  
  • Publications:  Pluznick, J. L, Rodriguez-Gil, D. J., Hull, M., Mistry, K., Gattone, V., Johnson, C. A., Weatherbee, S., Greer, C. A., Caplan, M. J. (2011) Renal cystic disease proteins play critical roles in the organization of the olfactory epithelium. PLOS One. 6(5):e19694. PMID: 21614130.
  • Bertuccio, C. A., Chapin, H. C., Cai, Y., Mistry, K., Chauvet, V., Somlo, S., Caplan, M. J. (2009). Polycystin-1 C-terminal cleavage is modulated by polycystin-2 expression. J Biol Chem. 284(31):21011-26. PMID:19491093
  • Takiar V, Mistry K, Carmosino M, Schaeren-Wiemers N, Caplan MJ. VIP17/MAL expression modulates epithelial cyst formation and ciliogenesis. Am J Physiol Cell Physiol. 2012 Oct 15;303(8):C862-71. doi: 10.1152/ajpcell.00338.2011. Epub 2012 Aug 15. PubMed PMID: 22895261; PubMed Central PMCID: PMC3469709.
  • Stoops EH, Hull M, Olesen C, Mistry K, Harder JL, Rivera-Molina F, Toomre D, Caplan MJ. The periciliary ring in polarized epithelial cells is a hot spot for delivery of the apical protein gp135. J Cell Biol. 2015 Oct 26;211(2):287-94. doi: 10.1083/jcb.201502045. PubMed PMID: 26504168; PubMed Central PMCID: PMC4621837.

Adele Ricciardi

  • Year: 7
  • Graduate Program: BME
  • Mentor Information: Mark Saltzman, BME
  • Education: U. of North Carolina, B.S., 2010 (Biology, Chemistry)
  • Publications: von Moeller H, Lerner R, Ricciardi A, Basquin C, Marsluff WF, Conti E. (2013). Structural and biochemical studies of SLIP1-SLBP identify DBP5 and eIF3g as SLIP1-binding proteins. Nucleic Acids Res. 1;41(16):7960-71. PMID 23804756.
  • Doyle F, Zaleski C, George AD, Stenson EK, Ricciardi A, Tenenbaum SA. (2008). Bioinformatic tools for studying post-transcriptional gene regulation: The UAlbany TUTR collection and other informatic resources. Methods Mol Biol. 419:39-52. PMID: 18369974.

Alexander Scherer

  • Year: 7
  • Graduate Program: Cell Biology
  • Mentor Information: Anthony Koleske, MB&B 
  • Research interest:  Abl family kinase regulation of actin cytoskeleton
  • Education: Stanford, B.S., 2011 (Chemistry)
  • A little about me:  I am originally from Indiana and enjoy playing softball and singing in the St. Thomas Episcopal Church Choir in my free time.

Samuel Sondalle

  • Year: 7
  • Graduate Program:  Genetics
  • Mentor Information: Susan Baserga, MB&B 
  • Research interest: My research focuses on investigating the molecular etiology of diseases due to defects in making ribosomes.
  • Education: Ripon College, B.A., 2011 (Biology, Chemistry)
  • A little about me:  I am a first-generation college student from the small town of Princeton, WI.  Outside of school I enjoy cooking and baking and playing piano.  I am active in the LGBTQ community at Yale and am happy to talk to any prospective LGBTQ students with questions about life at Yale. 
  • Publications:   Frey, B.L., Ladror, D.T., Sondalle, S.B., Krusemark, C.J., Jue, A.L., Coon, J.J., and Smith, L.M. (2013) Chemical derivatization of peptide carboxyl groups for highly efficient electron transfer dissociation. J Am Soc Mass Spectrom. Epub ahead of print. PMID: 23918461.
  • Zhou, Y., Cashman, T.J., Nevis, K.R., Obregon, P., Carney, S.A., Liu, Y., Gu, A., Mosimann, C., Sondalle, S., Peterson, R.E., Heideman, W., Burns, C.E., & Burns, C.G. (2011) Latent TGFβ binding protein 3 identifies a second heart field in zebrafish. Nature 474:645-648. PMID: 21623370.
  • Griffin, J.N., Sondalle, S.B., del Viso, F., Baserga, S.J., and Khokha, M.K. The ribosome biogenesis factor Nol11 is required for optimal rDNA transcription and craniofacial development in Xenopus. PLoS Genet (2015) 11(3): e1005018. doi:10.1371/journal.pgen.1005018. PMID: 25756904. Recommended by F1000Prime as being of special significance in its field for “interesting hypothesis” and “new finding.”
  • Sondalle, S.B., Baserga, S.J.*, Yelick, P.C.*. The contributions of the ribosome biogenesis protein, Utp5/WDR43, to craniofacial development . In press for Journal of Dental Research.  *co-corresponding authors

Alexander Svoronos

  • Year: 7
  • Graduate Program: MB&B
  • Mentor Information: Donald Engelman, MB&B 
  • Research interest:  Targeted drug delivery to cancer
  • Education: Brown, B.S., 2010 (Biomedical Engineering)
  • A little about me:  I was born and raised in Gainesville, FL. In my free time, I enjoy travel, hiking, yoyoing, and parkour. 
  • Publications:  Svoronos AA, Engelman DM, Slack FJ. Oncomir or Tumor Suppressor? The Duplicity of MicroRNAs in Cancer. Cancer Res. 2016 July 1; 76(13):3666-3670.
  • Cheng CJ, Bahal R, Babar IA, Pincus Z, Barrera F, Liu C, Svoronos A, Braddock DT, Glazer PM, Engelman DM, Saltzman WM, Slack FJ. MicroRNA silencing for cancer therapy targeted to the tumor microenvironment. Nature. 2015 Feb 5; 518:107-110. 
  • Svoronos AA, Tejavibulya N, Schell JY, Shenoy VB, Morgan JR. Micro-mold Design Controls the 3D Morphological Evolution of Self-assembling Multi-cellular Microtissues. Tissue Engineering Part A. 2014 April 9; 20(7-8):1134-1144. 
  • Wang H, Svoronos AA, Boudou T, Sakar MS, Schell JY, Morgan JR, Chen CS, Shenoy VB. Necking and failure of constrained 3D microtissues induced by cellular tension. Proc Natl Acad Sci. 2013 Dec 24; 110(52):20923-8.

Entering Class - 2010

Adrian Haimovich

Adrian Haimovich

  • Year:  8
  • Graduate Program: MCDB
  • Mentor Information: Farren Isaacs, MCDB 
  • Research interest: Synthetic and systems biology
  • Education:  Columbia, B.S., 2010 (Applied Mathematics)
  • A little about me:  I spent my pre-college years in central New Jersey, going to North Brunswick Township High School and learning to juggle a soccer ball with my left foot. I studied Applied Math at Columbia so I could try to get a grip on "big data" in the biomedical sciences. I became convinced that big data needed testable hypotheses (and ways to scale them), so after a stint at the NIH, I jumped into systems and synthetic biology. I am happy to call New Haven my home and my MD/PhD colleagues my friends. 
  • Publications: M Amiram, AD Haimovich, C Gang, YS Wang, HR Aerni, I Ntai, D Moonan, NJ Ma, AJ Rovner, SH Hong, N Kelleher, AL Goodman, MC Jewett, D Soll, J Rinehart, FJ Isaacs. “Evolution of translation machinery in recoded bacteria enables multi-site incorporation of nonstandard amino acids.” Nature Biotechnology. 2016, 33 (12), 1272-1279
  • AD Haimovich, P Muir, FJ Isaacs. “Genomes by Design”. Nature Reviews Genetics, 2015, 16 (9), 501-516. 
  • Pirman NL, Barber KW, Aerni HR, Ma NJ, Haimovich AD, Rogulina S, Isaacs FJ & Rinehart J “A flexible codon in genomically recoded Escherichia coli permits programmable protein phosphorylation.” Nat Comm. 2015, 6, 8130.
  • L Rover, AD Haimovich*, S Katz*, Z Li, MW Grome, B Gassaway, M Amiram, JR Patel, RR Gallagher, J Rinehart, FJ Isaacs. “Engineered organisms depend on a 21st amino acid for viability.” Nature, 2015, 518, 89-93. *Authors contributed equally to this work.
  • Sawyer, N., Gassaway, B.M., Haimovich, A.D., Isaacs, F.J., Rinehart, J., and Regan, L. Designed phosphoprotein recognition in Escherichia coli. ACS Chem. Biol., 2014, 9, 2502–2507.
  • SH Hong, I Ntai, AD Haimovich, N Kelleher, FJ Isaacs, M Jewett. "Cell-free protein synthesis from a release factor 1 deficient Escherichia coli activates efficient and multiple site-specific non-standard amino acid incorporation.” ACS Synth. Biol., 2013, Article ASAP.
  • M Lajoie, AJ Rovner, DB Goodman, HR Aerni, AD Haimovich, G Kuznetsov, JA Mercer, HH Wang, PA Carr, JA Mosberg, N Rohland, PG Schultz, JM Jacobson, J Rinehart, GM Church, FJ Isaacs. “Genomically Recoded Organisms Expand Biological Functions.” Science, 2013, Vol 342, 357-360.
  • AD Haimovich. “Methods, challenges, and promise of next-generation sequencing in cancer biology.” Yale Journal of Biol and Med, 2011, Vol 84 (4), 439.

Shihan Khan

  • Year:  8
  • Graduate Program:  BME
  • Mentor Information: Tarek Fahmy, BME
  • Education:  University of Michigan, B.S., 2010 (BME)

Rebecca Liu

  • Year:  8
  • Graduate Program:   Immunobiology
  • Mentor Information: Jordan Pober, Immunobiology 
  • Research interest:  Human immunology (Inflammation; Transplantation; Autoimmunity)
  • Education:  Duke University, B.S., 2010 (Biology)
  • A little about me:  Outside of the lab and hospital, I enjoy going on culinary adventures, exploring different restaurants in New England and around the world. My other hobbies include photography, hiking, and golf. In addition, I am passionate about health education and community outreach, and have been actively involved in mentoring high school/undergraduate students who are interested in a career in science and medicine.
  • Publication:  Jane-Wit D, Surovtseva YV, Qin L, Li G, Liu R, Clark P, Manes TD, Wang C, Kashgarian M, Kirkiles-Smith NC, Tellides G, Pober JS, Complement membrane attack complexes activate noncanonical NF-κB by forming an Akt+NIK+ signalosome on Rab5+ endosomes. Proc Natl Acad Sci. 2015 Jul 20. PMID: 26195760
  • Abrahimi P, Liu R, and Pober JS, Blood Vessels in Allotransplantation, Am J Transplant. 2015 Jul; 15(7):1748-54. PMID: 25807965
  • Yao Y, Liu R, Shin MS, Trentalange M, Allore H, Nassar A, Kang I, Pober J, and Montgomery RR, CyTOF supports efficient detection of immune cell subsets from small samples. Journal of Immunological Methods, 2014. 415: p. 1-5. PMID: 25450003.
  • Nair SK, De Leon G, Boczkowski D, Schmittling R, Xie W, Staats J, Liu R, Johnson LA, Weinhold K, Archer GE, Sampson JH, Mitchell DA, Recognition and killing of autologous, primary glioblastoma tumor cells by human cytomegalovirus pp65-specific cytotoxic T cells. Clinical Cancer Research, 2014. 20(10): p. 2684-94. PMCID: PMC4059212.
  • Liu R, Mitchell DA, Survivin as an immunotherapeutic target for adult and pediatric malignant brain tumors. Cancer Immunology Immunotherapy, 2010. 59(2): p. 183-93. PMID: 19756596.

Kelsey Loeliger

  • Year: 8
  • Graduate Program:   Public Health
  • Mentor Information: Christian Tschudi, Public Health 
  • Research interest:  Epidemiology of Microbial Diseases, The impact of incarceration and release from a correctional facility on linkage to care, retention in care, and viral suppression among people living with HIV in Connecticut (Christian Tschudi, PhD; Frederick Altice, MD)
  • Education:  University of Maryland, Baltimore County (UMBC), B.S., 2008 (Biology)
  • A little about me:  My research interests broadly include infectious diseases, substance use disorders, women’s health, mental health, interpersonal violence, international health, and social justice. Affiliated with the Yale School of Public Health’s Epidemiology of Microbial Diseases Department, I am interested in conducting clinical and community-based research with an emphasis on marginalized populations and the social constructs surrounding infectious diseases. My work is aimed at developing a better understanding of the many interconnected factors that play a role in the prevention, progression, transmission, and treatment of these diseases. My previous research with Drs. Gerald Friedland and Sheela Shenoi in South Africa explored the personal and systematic barriers HIV patients face with regard to initiating and adhering to antiretroviral therapy. Under the direction of Dr. Frederick Altice, areas of study have also included women with substance use disorders in Malaysia and how the interplay between substance abuse, HIV/AIDS, mental health disorders, and violence as well as other social circumstances impact the health of the individual. My current dissertation research aims to track and improve HIV treatment outcomes in Connecticut criminal justice populations after release from prison or jail. As an undergraduate, I studied the retroviral mechanism for genome packaging in the Howard Hughes Medical Institute biochemistry laboratory of Michael Summers, PhD. After graduating, I worked as a post-baccalaureate research fellow (IRTA) at the National Heart Lung and Blood Institute where I conducted translational research on the role of chronic inflammation in bone marrow failure. 
  • Publication:  Kelsey B. Loeliger, Mary L. Biggs, David Seal, Michael S. Gordon, Curt G. Beckwith, Irene Kuo, Jeremy D. Young, Carol E. Golin, Redonna Chandler, Wendee M. Wechsberg, Frederick L. Altice, William Cunningham, Sandra A. Springer. Gender Differences in HIV Risk Behaviors Among HIV-Infected and Uninfected Persons Involved in the U.S. Criminal Justice System: A “Seek Test Treat Retain” Harmonization Project. (Manuscript in preparation for JAIDS.)
  • Loeliger K.B., Niccolai, L.M., Mtungwa, N., Moll, A., Shenoi, S. “I have to push him with a wheelbarrow to the clinic”: Community health workers’ roles, needs, and strategies to improve HIV care in rural South Africa. (Manuscript submitted 26 April 2016 to AIDS Patient Care & STDs.)
  • Loeliger, K.B., Niccolai, L.M., Mtungwa, L.N., Moll, A., & Shenoi, S.V. (2016). Antiretroviral therapy initiation and adherence in rural South Africa: community health workers' perspectives on barriers and facilitators. AIDS Care, 1-12. doi:10.1080/09540121.2016.1164292
  • Loeliger K.B., Marcus R., Wickersham J.A., Pillai, V., Kamarulzaman A., Altice F.L. (2015) The syndemic of HIV, HIV-related risk and multiple co-morbidities among women who use drugs in Malaysia: Important targets for intervention. Addict Behav, 53: 31-39. doi: 10.1016/j.addbeh.2015.09.013
  • Wickersham J.A., Loeliger K.B., Marcus R., Pillai, V., Kamarulzaman A., Altice F.L. (2015) Patterns of Substance Use and Correlates of Active and Remote Injection Drug Use Among Women in Malaysia. American Journal of Drug and Alcohol Abuse. 2015 Dec 4:1-13. doi: 10.3109/00952990.2015.1101467
  •  Al-Darraji HA, Wong KC, Yeow DG, Fu JJ, Loeliger K, Paiji C, Kamarulzaman A, Altice FL, 2014, Tuberculosis screening in a novel substance abuse treatment center in Malaysia: implications for a comprehensive approach for integrated care. Journal of substance abuse treatment, 46(2):144-9. PMID: 24074846 
  • Zhao YO, Kurscheid S, Zhang Y, Liu L, Zhang L, Loeliger K, Fikrig E, 2012, Enhanced survival of Plasmodium-infected mosquitoes during starvation. PloS one, 7(7):e40556. PMID: 22808193; PMCID: PMC3393683

Goran Micevic

  • Year:  8
  • Graduate Program:  Experimental Pathology
  • Mentor Information: Marcus Bosenberg, Pathology
  • Education:  Iowa State University, B.S., 2010 (Biochemistry)

Laura Pappalardo

  • Year: 8
  • Graduate Program: INP
  • Mentor Information: Stephen Waxman, Neurobiology 
  • Research interest: Molecular mechanisms by which neuroglia respond to and regulate CNS injury and disease. Specifically, I am examining the contribution of voltage-gated sodium channels to astrogliosis, with implication for multiple sclerosis and spinal cord injury.
  • Education: University of Texas, Pharm.D., 2010  (Pharmacy)
  • A little about me: I’m a displaced Texan, but after 6 years on the East Coast and marrying a New Yorker, it’s starting to feel like home. My husband works in NYC so we’re currently living the commuter lifestyle in Stamford, CT, and while podcasts are a must, it’s certainly feasible. When I’m not at lab, I spend time traveling, trying to hunt down decent Mexican food (no luck), and devising a method to diet my obese cat (also no luck). 
  • Publications: Pappalardo LW, Black JA, Waxman SG. Sodium channels in neuroglia (2016) Glia.
  • Pappalardo LW, Black JA, Waxman SG. Dynamics of sodium channel Nav1.5 expression in astrocytes in mouse models of multiple sclerosis (2014) NeuroReport; 25: 1208-1215.
  • Pappalardo LW, Samad OA, Black JA, Waxman SG. Voltage-gated sodium channel Nav1.5 contributes to astrogliosis in an in vitro model of glial injury via reverse Na+ /Ca2+ exchange (2014) Glia; 62: 1162-1175.

Max Petersen

  • Year:  8
  • Graduate Program:    C&M Physiology
  • Mentor Information: Gerald Shulman, C&M Physiology 
  • Research interest: Type 2 Diabetes
  • Education:  Augustana College, B.A., 2010 (Biochemistry, Pre-Med & Religious Studies)
  • A little about me:  I love music, working out, and hanging out with my Golden Retriever Charlie. 
  • Publication: Petersen MC, Jurczak MJ. CrossTalk opposing view: Intramyocellular ceramide accumulation does not modulate insulin resistance. 2016. Journal of Physiology (Epub ahead of print) PMID: 26997581.
  • Camporez JP, Petersen MC, Abulizi A, Moreira GV, Jurczak MJ, Friedman G, Haqq CM, Petersen KF, Shulman GI. Anti-myostatin antibody increases muscle mass and strength and improves insulin sensitivity in old mice. 2016. Proceedings of the National Academy of Sciences of
    the United States of America 113(8):2212-7. PMID: 26858428
  • Camporez JP, Jornayvaz FR, Petersen MC, Pesta D, Guigni BA, Serr J, Zhang D, Kahn M, Samuel VT, Jurczak MJ, Shulman GI. (2013) Cellular Mechanisms by Which FGF21 Improves Insulin Sensitivity in Male Mice. Endocrinology 154:3099-109. PMID: 23766126; PMCID: PMC3749479

Susan Scanlon

  • Year:  8
  • Graduate Program:   Experimental Pathology
  • Mentor Information: Peter Glazer, Genetics 
  • Research interest:  Hypoxia and DNA repair
  • Education:  Yale University, B.S., 2010 (MB&B & Physics)
  • A little about me:  Outside of the hospital and the lab, I mainly spend time with family and friends and try to stay active. I enjoy hiking, swimming, biking, frisbee, golf, and crossfit. I also like apple picking, eating ice cream, the Red Sox and Patriots, listening to music, doing puzzles, and playing Scrabble. 
  • Publication:  Scanlon, S.E. and P.M. Glazer. (2013). Genetic Instability Induced by Hypoxic Stress. In D. Mittelman (Ed.), Stress-Induced Mutagenesis (pp. 151-182). New York: Springer.
  • Glazer PM, Hegan DC, Lu Y, Czochor J, Scanlon SE, 2013, Hypoxia and DNA repair. The Yale journal of biology and medicine, 86(4):443-51. PMID: 24348208; PMCID: PMC3848098
  • Scanlon, S.E. and P.M. Glazer. Multifaceted control of DNA repair pathways by the hypoxic tumor microenvironment. DNA Repair (Amst). Aug 2015;32:180-9. PMCID: PMC4522377.
  • Scanlon, S.E. and P.M. Glazer. Hypoxic Stress Facilitates Acute Activation and Chronic Down-Regulation of Fanconi Anemia Proteins. Molecular Cancer Research. Jul 2014;.12(7):1016-28. PMCID: PMC4101147.

Alexandra Thomas

  • Year: 8
  • Graduate Program:   INP
  • Mentor Information: Ronald Duman, Neurobiology
  • Education:  Princeton University, B.A., 2009 (Public & International Affairs, Neuroscience)

Wendy Xiao

  • Year:  8
  • Graduate Program:   INP
  • Mentor Information:  Hal Blumenfeld, Neurobiology
  • Education:  Emory University, B.S., 2009 (Anthropology & Neuroscience)

Genevieve Yang

  • Year:  8
  • Graduate Program:   INP
  • Mentor Information: Alan Anticevic, Psychiatry
  • Education:  Columbia University, B.A., 2008 (Biology)

Ke Zhang

  • Year:  8
  • Graduate Program:   Cell Biology
  • Mentor Information: Anthony Koleske, MB&B
  • Education:  MIT, B.S., 2010 ( Chemistry & Physics)

Entering Class - 2009

Alexander Bazazi

  • Year: 9
  • Graduate Program:   Epidemiology of Microbial Diseases
  • Mentor Information: Robert Heimer, Public Health
  • Education:  Boston University, B.A., 2008 (Philosophy) 
  • Publications: Bazazi A.R., Zelenev A.Z., Altice F.L. (2013) Letter to the Editor: regarding "Individual and Neighborhood Correlates of Membership in Drug-Using Networks With a Higher Prevalence of HIV in New York City (2006-2009)". Ann Epidemiology.  PMID: 23972897.
  • Lim SH, Bazazi AR, Sim C, Choo M, Altice FL, Kamarulzaman A. High rates of unprotected anal intercourse with regular and casual partners and associated risk factors in a sample of ethnic Malay men who have sex with men (MSM) in Penang, Malaysia. Sexually Transmitted Infections. [In Press]
  • Fu JJ, Bazazi AR, Altice FL, Mohamed MN, Kamarulzaman A. Absence of antiretroviral therapy and other risk factors for morbidity and mortality in Malaysian compulsory drug detention and rehabilitation centers. PLoS ONE 2012; 7(9): e44249. PMCID: PMC3445567.
  • Bachireddy C, Bazazi AR, Kavasery R, Govindasamy S, Kamarulzaman A, Altice FL. Attitudes toward opioid substitution therapy and pre-incarceration HIV transmission behaviors among HIV-infected prisoners in Malaysia: Implications for secondary prevention. Drug and Alcohol Dependence 2011;116(1-3):151-157. PMID: 21232882.
  • Fu JJ, Bazazi AR, Altice FL. Preserving human rights in the era of ‘test and treat’ for HIV prevention. American Journal of Public Health 2011;101(2):198-199. PMID: 21228278.

Simon Gray

  • Year: 9
  • Graduate Program:  Immunobiology
  • Mentor Information:  Susan Kaech, Immunobiology 
  • Research interest: My thesis research in the laboratory of Susan Kaech focuses on understanding how epigenetic mechanisms regulate the commitment or plasticity of antiviral CD8+ T cell populations.
  • Education:  University of Maryland, B.S., 2008 (Chemical Engineering); University of Cambridge, M.O., 2009 (Chemical Engineering) 
  • A little about me: In my spare time I enjoy swimming, cycling, triathlons, sailing, kite-surfing, beekeeping and playing the piano. 
  • Publication: Gray SM, Bloch MH, 2012, Systematic review of proinflammatory cytokines in obsessive-compulsive disorder. Current psychiatry reports, 14(3):220-8. PMID: 22477442; PMCID: PMC3625952 
  • Gray S, Bloch M. Systematic Review of Proinflammatory Cytokines in Obsessive-Compulsive Disorder. Curr Psychiatry Rep 2012;14:220-8
  • Gray SM, Kaech SM, Staron MM. The interface between transcriptional and epigenetic control of effector and memory CD8(+) T-cell differentiation. Immunol Rev 2014;261:157-68.
  • Staron MM, Gray SM, Marshall HD, et al. The transcription factor FoxO1 sustains expression of the inhibitory receptor PD-1 and survival of antiviral CD8(+) T cells during chronic infection. Immunity 2014;41:802-14.
  • Cui G, Staron MM, Gray SM, et al. IL-7-Induced Glycerol Transport and TAG Synthesis Promotes Memory CD8 T Cell Longevity. Cell 2015;161:750-61.
  • Laidlaw BJ, Cui W, Amezquita RA, Gray SM, Guan T, Lu Y, Kleinstein SH, Craft J, Kaech SM. IL-10 Production by Regulatory CD4+ T Cells During Resolution of Infection Promotes Memory CD8+ T Cell Maturation. Nature Immunology 2015;16:871-879.
  • Ray JP, Staron MM, Shyer JA, Ho PC, Gray SM, Laidlaw BJ, Araki K, Ahmed R, Kaech SM, Craft J. The Interleukin-2-mTORc1 Kinase Axis Defines the Signaling, Differentiation, and Metabolism of T Helper 1 and Follicular B Helper T Cells. Immunity 2015;43(4): 690-702.

Marco Ramos

  • Year: 9
  • Graduate Program:  History of the Science of Medicine
  • Mentor Information: John Warner, History of Medicine
  • Education:  Columbia University, B.A., 2007 (Pre-Med) 
  • Publication: Ramos MA, 2013, Drugs in context: a historical perspective on theories of psychopharmaceutical efficacy. The Journal of nervous and mental disease, 201(11):926-33. PMID: 24177478 
  • Ramos MA, 2013, Psychiatry, authoritarianism, and revolution: the politics of mental illness during military dictatorships in Argentina, 1966-1983. Bulletin of the history of medicine, 87(2):250-78. PMID: 23811712

Andrew H. Smith

  • Year: 9
  • Graduate Program:  INP
  • Mentor Information:  Joel Gelernter, Neurobiology 
  • Research interest:  Neuroscience, genetics, addiction
  • Education:  Northwestern, B.A., 2004 (American Studies) 
  • A little about me: From Shaker Heights, OH. Enrolled in the Interdepartmental Neuroscience Program.
  • Publication: Jensen KP, Smith AH, Herman AI, Farrer LA, Kranzler HR, Sofuoglu M, Gelernter J. A protocadherin gene cluster regulatory variant is associated with nicotine withdrawal and the urge to smoke. Molecular Psychiatry, 2016.
  • Polimanti R, Zhang H, Smith AH, Zhao H, Farrer LA, Kranzler HR, Gelernter J. Genome-wide association study of body mass index in subjects with alcohol dependence. Addiction Biology, 2015.
  • Clarke TK, Smith AH, Gelernter J, Kranzler HR, Farrer, LA, Hall LS, Fernandez-Pujals AM, MacIntyre DJ, Smith BH, Hocking LJ, Padmanabhan S, Hayward C, Thomson PA, Porteous DJ, Deary IJ, McIntosh AM. Polygenic risk for alcohol dependence associates with alcohol consumption, cognitive function and social deprivation in a population-based cohort. Addiction Biology, 2015.
  • Niculescu A, Levey D, Le-Niculescu H, Frank J, Ayalew M, Jain N, Kirlin B, Learman R, Winiger E, Rodd Z, Shekhar A, Schork N, Kiefe F, Wodarz N, Müller-Myhsok B, Dahmen N, Nöthen M, Sherva R, Farrer L, Smith AH, Kranzler H, Rietschel M, and Gelernter J. Genetic Risk Prediction and Neurobiological Understanding of Alcoholism. Translational Psychiatry, 2014.
  • Gelernter J, Kranzler HR, Sherva R, Almasy L, Koesterer R, Smith AH, Anton R, Preuss UW, Ridinger M, Rujescu D, Wodarz N, Zill P, Han S, Zhao H, Farrer LA. Genomewide association study of alcohol dependence: Significant findings in African- and European-Americans including novel loci. Molecular Psychiatry, 2014.

Eleanor Thomas

  • Year: 9
  • Graduate Program:  INP
  • Mentor Information:  Arthur Horwich, Genetics
  • Education:  Brown University, B.A., B.S. 2007 (Classics, Neuroscience)

Entering Class - 2008

Alexandria Marino

  • Year:  10
  • Graduate Program: INP
  • Mentor Information: James Mazer, Neurobiology 
  • Research interest: I study how we understand navigate the visual world but examining responses of visual neurons recorded from monkeys that are trained to do complex behavioral tasks.
  • Education:Yale University, B.A., 2005 (Cognitive Science)
  • A little about me:  I am originally from West Virginia and spent a few years teaching high school chemistry in Los Angeles before joining the program. When I'm not in lab, I'm likely knitting, playing violin, or hanging out with my husband and two cats. 
  • Publications:  Golomb, J.D., Marino, A.C., Chun, M.M., and Mazer, J.A. 2011 Attention doesn't slide: Spatiotopic updating after eye movements instantiates a new, discrete attentional locus. Attention, Perception, and Psychophysics. 73(1): 7-14. PMID: 21258903, PMCID: PMC3097429
  • van Erp TG, Thompson PM, Kieseppä T, Bearden CE, Marino AC, Hoftman GD, Haukka J, Partonen T, Huttunen M, Kaprio J, Lönnqvist J, Poutanen VP, Toga AW, Cannon TD. Hippocampal morphology in lithium and non-lithium-treated bipolar I disorder patients, non-bipolar co-twins, and control twins. Hum Brain Mapp. 2012 Mar;33(3):501-10. doi: 10.1002/hbm.21239. Epub 2011 Mar 31. PMID: 21455943
  • Marino, A. C., and Mazer, J.A.. (2016). Perisaccadic updating of visual representations and attentional states: Linking behavior and neurophysiology. Frontiers Systems Neuroscience. Feb 5;10:3.

Curtis Perry

  • Year:  10
  • Graduate Program: Immunobiology
  • Mentor Information: Susan Kaech, Immunobiology
  • Education:Yale University, B.S., 2007 (Biochemistry)