Gordon M Shepherd MD, DPhil

Professor of Neurobiology

Departments & Organizations

Yale Combined Program in the Biological and Biomedical Sciences (BBS): Computational Biology and Bioinformatics | Neuroscience

Office of Student ResearchInterdepartmental Neuroscience ProgramNeurobiologyShepherd Lab

Research Interests

Neuronal Dendrites; Dendritic Spines; Synaptic Organization; Olfactory System; Brain Microcircuits; Computational Neuroscience; Neuroinformatics; Functional Connectomes more...


  • B.S., Iowa State College, 1955
  • M.D., Harvard University, 1959
  • D.Phil., Oxford University, 1962

Selected Publications

  • Sparse coding and lateral inhibition arising from balanced and unbalanced dendrodendritic excitation and inhibition. Yu Y, Migliore M, Hines ML, Shepherd GM. J Neurosci. 2014 Oct 8;34(41):13701-13. doi: 10.1523/JNEUROSCI.1834-14.2014.
  • Distributed organization of a brain microcircuit analyzed by three-dimensional modeling: the olfactory bulb. Migliore M, Cavarretta F, Hines ML, Shepherd GM. Front Comput Neurosci. 2014 Apr 29;8:50. doi: 10.3389/fncom.2014.00050. eCollection 2014.
  • Shepherd GM. Ann N Y Acad Sci. 2009 Jul;1170:215-23. No abstract available. PMID: 19686140 [PubMed - in process]



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