Biff Forbush PhD

Professor of Cellular and Molecular Physiology

Departments & Organizations

Yale Combined Program in the Biological and Biomedical Sciences (BBS): Molecular Medicine, Pharmacology, and Physiology

Office of Student ResearchCellular & Molecular Physiology: Membrane Proteins - Pumps and Transporters; Epithelial Transport of Ions and Solutes; Physiology and Integrative Medical Biology Track; Graduate Program in Cellular and Molecular Physiology

Research Interests

Membrane Proteins; Epithelial Transport; Ion Transport; Na-K-Cl Cotransporter; Nerve Cells; Muscle Cells more...


  • Ph.D., Johns Hopkins University, 1975

Selected Publications

  • Gimenez I and Forbush B. The residues determining differences in ion affinities among the alternative splice variants F, A, and B of the mammalian renal Na-K-Cl cotransporter (NKCC2). J Biol Chem, 282:6540-6547, 2007.
  • Dowd BF and Forbush B. PASK (proline-alanine-rich STE20-related kinase), a regulatory kinase of the Na-K-Cl cotransporter (NKCC1). J Biol Chem, 278:27347-53, 2003.



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